Abstract In the theory of protoplanetary disk turbulence, a widely adopted ansatz, or assumption, is that the turnover frequency of the largest turbulent eddy, ΩL, is the local Keplerian frequency ΩK. In terms of the standard dimensionless Shakura–Sunyaevαparameter that quantifies turbulent viscosity or diffusivity, this assumption leads to characteristic length and velocity scales given respectively by and , in whichHandcare the local gas scale height and sound speed. However, this assumption is not applicable in cases when turbulence is forced numerically or driven by some natural processes such as vertical shear instability. Here, we explore the more general case where ΩL≥ ΩKand show that, under these conditions, the characteristic length and velocity scales are respectively and , where is twice the Rossby number. It follows that , where is the root-mean-square average of the turbulent velocities. Properly allowing for this effect naturally explains the reduced particle scale heights produced in shearing box simulations of particles in forced turbulence, and it may help with interpreting recent edge-on disk observations; more general implications for observations are also presented. For , the effective particle Stokes numbers are increased, which has implications for particle collision dynamics and growth, as well as for planetesimal formation. 
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                            Emergence of Escherichia coli critically buckled motile helices under stress
                        
                    
    
            Bacteria under external stress can reveal unexpected emergent phenotypes. We show that the intensely studied bacteriumEscherichia colican transform into long, highly motile helical filaments poized at a torsional buckling criticality when exposed to minimum inhibitory concentrations of several antibiotics. While the highly motile helices are physically either right- or left-handed, the motile helices always rotate with a right-handed angular velocity , which points in the same direction as the translational velocity of the helix. Furthermore, these helical cells do not swim by a “run and tumble” but rather synchronously flip their spin and thus translational velocity—backing up rather than tumbling. By increasing the translational persistence length, these dynamics give rise to an effective diffusion coefficient up to 20 times that of a normalE. colicell. Finally, we propose an evolutionary mechanism for this phenotype’s emergence whereby the increased effective diffusivity provides a fitness advantage in allowing filamentous cells to more readily escape regions of high external stress. 
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                            - Award ID(s):
- 1734030
- PAR ID:
- 10080539
- Publisher / Repository:
- Proceedings of the National Academy of Sciences
- Date Published:
- Journal Name:
- Proceedings of the National Academy of Sciences
- Volume:
- 115
- Issue:
- 51
- ISSN:
- 0027-8424
- Page Range / eLocation ID:
- p. 12979-12984
- Format(s):
- Medium: X
- Sponsoring Org:
- National Science Foundation
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