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			<titleStmt><title level='a'>Gas-rich, Field Ultra-diffuse Galaxies Host Few Gobular Clusters</title></titleStmt>
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				<publisher></publisher>
				<date>12/28/2022</date>
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				<bibl> 
					<idno type="par_id">10434055</idno>
					<idno type="doi">10.3847/2041-8213/acaaab</idno>
					<title level='j'>The Astrophysical Journal Letters</title>
<idno>2041-8205</idno>
<biblScope unit="volume">942</biblScope>
<biblScope unit="issue">1</biblScope>					

					<author>Michael G. Jones</author><author>Ananthan Karunakaran</author><author>Paul Bennet</author><author>David J. Sand</author><author>Kristine Spekkens</author><author>Burçin Mutlu-Pakdil</author><author>Denija Crnojević</author><author>Steven Janowiecki</author><author>Lukas Leisman</author><author>Catherine E. Fielder</author>
				</bibl>
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			<abstract><ab><![CDATA[Abstract                          We present Hubble Space Telescope imaging of 14 gas-rich, low-surface-brightness galaxies in the field at distances of 25–36 Mpc, with mean effective radii and              g              -band central surface brightnesses of 1.9 kpc and 24.2 mag arcsec              −2              . Nine meet the standard criteria to be considered ultra-diffuse galaxies (UDGs). An inspection of point-like sources brighter than the turnover magnitude of the globular cluster luminosity function and within twice the half-light radii of each galaxy reveals that, unlike those in denser environments, gas-rich, field UDGs host very few old globular clusters (GCs). Most of the targets (nine) have zero candidate GCs, with the remainder having one or two candidates each. These findings are broadly consistent with expectations for normal dwarf galaxies of similar stellar mass. This rules out gas-rich, field UDGs as potential progenitors of the GC-rich UDGs that are typically found in galaxy clusters. However, some in galaxy groups may be directly accreted from the field. In line with other recent results, this strongly suggests that there must be at least two distinct formation pathways for UDGs, and that this subpopulation is simply an extreme low surface brightness extension of the underlying dwarf galaxy population. The root cause of their diffuse stellar distributions remains unclear, but the formation mechanism appears to only impact the distribution of stars (and potentially dark matter), without strongly impacting the distribution of neutral gas, the overall stellar mass, or the number of GCs.]]></ab></abstract>
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<div xmlns="http://www.tei-c.org/ns/1.0"><head n="1.">Introduction</head><p>The study of low-surface-brightness (LSB) galaxies, including dwarf galaxies, has a long history <ref type="bibr">(Sandage &amp; Binggeli 1984;</ref><ref type="bibr">Impey et al. 1988;</ref><ref type="bibr">Thompson &amp; Gregory 1993;</ref><ref type="bibr">Jerjen et al. 2000;</ref><ref type="bibr">Conselice et al. 2003;</ref><ref type="bibr">Mieske et al. 2007</ref>). However, the abundance of the most extreme LSB galaxies was not fully appreciated until relatively recently when hundreds of ultra-diffuse galaxies (UDGs) were identified in nearby galaxy clusters <ref type="bibr">(Koda et al. 2015;</ref><ref type="bibr">Mihos et al. 2015</ref>; <ref type="bibr">van Dokkum et al. 2015;</ref><ref type="bibr">Yagi et al. 2016;</ref><ref type="bibr">van der Burg et al. 2016;</ref><ref type="bibr">Wittmann et al. 2017;</ref><ref type="bibr">Venhola et al. 2017;</ref><ref type="bibr">Zaritsky et al. 2019)</ref>. Soon large samples of UDGs were identified in all environments from those in clusters, to galaxy groups <ref type="bibr">(Trujillo et al. 2017;</ref><ref type="bibr">van der Burg et al. 2017;</ref><ref type="bibr">Roman &amp; Trujillo 2017;</ref><ref type="bibr">Bennet et al. 2018;</ref><ref type="bibr">Spekkens &amp; Karunakaran 2018)</ref>, and to the field <ref type="bibr">(Leisman et al. 2017;</ref><ref type="bibr">Janowiecki et al. 2019;</ref><ref type="bibr">Prole et al. 2019;</ref><ref type="bibr">Roman et al. 2019;</ref><ref type="bibr">Karunakaran et al. 2020)</ref>. These findings led to a number of potential formation models including both internal mechanisms (e.g., <ref type="bibr">Amorisco &amp; Loeb 2016;</ref><ref type="bibr">Di Cintio et al. 2017;</ref><ref type="bibr">Chan et al. 2018</ref>) and those relying on external, environmental effects <ref type="bibr">(Conselice 2018;</ref><ref type="bibr">Carleton et al. 2019;</ref><ref type="bibr">Tremmel et al. 2020</ref>). It has also been hypothesized that UDGs may be the result of a combination of internal and external mechanisms, either acting jointly <ref type="bibr">(Martin et al. 2019;</ref><ref type="bibr">Jackson et al. 2021)</ref> or with different pathways being responsible for different subsets of the UDG population <ref type="bibr">(Papastergis et al. 2017;</ref><ref type="bibr">Pandya et al. 2018;</ref><ref type="bibr">Jiang et al. 2019;</ref><ref type="bibr">Liao et al. 2019;</ref><ref type="bibr">Wright et al. 2021;</ref><ref type="bibr">Buzzo et al. 2022)</ref>. These mechanisms range from early truncated growth, star formation feedback, intrinsic halo properties, tidal rarification, and mergers (for a recent summary of these proposed mechanisms, see <ref type="bibr">Jones et al. 2021)</ref>. While a number of works have argued that UDGs may have multiple formation pathways, it is still unclear whether UDGs in the field are directly related to those in denser environments. Could field UDGs be the progenitors of UDGs in denser environments, or are these largely distinct populations that formed via unrelated processes?</p><p>A key metric related to the early stages of formation of a galaxy is the number of old globular clusters (GCs) that it hosts. The richness of a galaxy's GC system is strongly correlated with its total mass (e.g., <ref type="bibr">Blakeslee et al. 1997;</ref><ref type="bibr">Harris et al. 2013;</ref><ref type="bibr">Zaritsky 2022)</ref>, for which dark matter (DM) halo mass, stellar masses, and luminosity (in order of decreasing linearity and tightness of the relation) may all be used as proxies. GCs therefore offer a means to probe the DM halo masses of UDGs (which appear to follow the established relation; <ref type="bibr">Harris et al. 2017</ref>), but also a means to compare subsets of the UDG population that are otherwise similar in terms of their surface brightness and stellar mass. Some investigations of the GC systems of UDGs in clusters have found them to be extraordinarily rich <ref type="bibr">(Beasley et al. 2016;</ref><ref type="bibr">Beasley &amp; Trujillo 2016;</ref><ref type="bibr">Dokkum et al. 2016;</ref><ref type="bibr">Peng &amp; Lim 2016</ref>; <ref type="bibr">van Dokkum et al. 2017)</ref>, while others have argued their GC systems are less extreme <ref type="bibr">(Amorisco et al. 2018;</ref><ref type="bibr">Forbes et al. 2020;</ref><ref type="bibr">Lim et al. 2020;</ref><ref type="bibr">Somalwar et al. 2020;</ref><ref type="bibr">Saifollahi et al. 2021)</ref>. However, even with the lower GC count estimates, UDGs still host richer GC systems on average than other dwarf galaxies of equivalent luminosity or stellar mass, but they correspond to dwarf-mass DM halos (e.g., 10 10 -10 11.5 M e ), not Milky Way-mass halos (e.g., &#8764;10 12 M e ). <ref type="bibr">Jones et al. (2021)</ref> used Hubble Space Telescope (HST) observations to identify GC candidates (GCCs) in two group UDGs that appeared to have (tidal) stellar streams connecting them to their respective hosts <ref type="bibr">(Bennet et al. 2018)</ref>. Unlike most UDGs in clusters, these UDGs appeared to host a small number of GCs, roughly in line with expectations for typical dwarf galaxies. This strongly suggested that these were previously regular dwarfs that were "puffed up" by tidal interactions with their hosts, after falling into a group (e.g., <ref type="bibr">Carleton et al. 2019)</ref>. This would make them distinct from cluster UDGs, but also UDGs that became ultra-diffuse while in the field, presumably via some internal mechanism. However, a significant caveat to this finding still remains: we are still largely ignorant of the properties of the GC systems of field UDGs. They may also be consistent with those of typical dwarf galaxies, in which case it would be less clear whether group UDGs such as those identified by <ref type="bibr">Bennet et al. (2018)</ref> are truly distinct from those in the field. They plausibly could have already been ultra-diffuse prior to falling into their current groups, and the evidence of tidal interactions may have no bearing on their status as UDGs. Equally, it may be possible that field UDGs are instead GC-rich and represent the progenitors of UDGs in cluster environments.</p><p>In this work we address this missing information by performing a census of the GCCs in 14 UDGs and LSB galaxies in the field, using HST Wide Field Camera 3 (WFC3) snapshot observations. The paper is organized as follows. In Section 2 we describe our target sample and observational strategy. In Section 3 we explain our approach to selecting GCCs and present the resulting GC counts. In Section 4 we discuss the interpretation of these results, and present our conclusions in Section 5.</p></div>
<div xmlns="http://www.tei-c.org/ns/1.0"><head n="2.">Sample and Observations</head><p>The UDGs targeted in this work are drawn from the H I- bearing UDGs sample of <ref type="bibr">Janowiecki et al. (2019)</ref>. These are UDGs originally detected through their H I line emission in the Arecibo Legacy Fast ALFA (Arecibo L-band Feed Array) survey (ALFALFA; <ref type="bibr">Giovanelli et al. 2005;</ref><ref type="bibr">Haynes et al. 2011</ref><ref type="bibr">Haynes et al. , 2018) )</ref> and subsequently identified as UDGs based on their LSB counterparts in Sloan Digital Sky Survey (SDSS; <ref type="bibr">York et al. 2000)</ref> images. This catalog is an expansion and revision of the initial ALFALFA-based catalog of <ref type="bibr">Leisman et al. (2017)</ref>. Part of this revision was to drop the explicit requirement for objects to be isolated from other, larger galaxies (as was the case in <ref type="bibr">Leisman et al. 2017)</ref>, but most of the expansion of the sample is simply the result of including more of the full footprint of ALFALFA, which was unavailable when the original catalog was published.</p><p>Despite no explicit requirement for isolation, <ref type="bibr">Janowiecki et al. (2019)</ref> demonstrate that H I-bearing UDGs reside in environments typical of other similar-mass, gas-rich galaxies in the ALFALFA survey, which are generally low-mass centrals in their own halos <ref type="bibr">(Guo et al. 2017)</ref>. Furthermore, none of our targets were matched to known groups by <ref type="bibr">Jones et al. (2020)</ref>. Thus, these H I-bearing UDGs are bona fide field objects, not satellites of larger galaxies or groups. This distinguishes this sample from other field UDG samples, such as those identified in the Mass Assembly of early Type gaLAxies with their fine Structures (MATLAS) survey <ref type="bibr">(Habas et al. 2020;</ref><ref type="bibr">Marleau et al. 2021)</ref>, whose members, though in low-density environments, are still mostly satellites.</p><p>The detection of H I is a prerequisite for the identification of these objects, meaning that, unlike most UDG samples, which are based purely on photometry, all have known spectroscopic redshifts. Furthermore, the parent sample <ref type="bibr">(Janowiecki et al. 2019)</ref> does not consider candidates within &#8764;25 Mpc, which removes those with the largest fractional uncertainties (due to peculiar velocities) on their redshift-based distance estimates. A maximum distance limit of 120 Mpc was also applied, as beyond this limit the projected distance corresponding to Arecibo's &#8764;3 5 beam can complicate the robust identification of the optical counterparts of H I detections, especially for LSB objects.</p><p>The UDGs and LSB galaxies in our observed sample are shown in Figure <ref type="figure">1</ref> relative to UDGs in a cluster environment.</p></div>
<div xmlns="http://www.tei-c.org/ns/1.0"><head n="2.1.">Hubble Space Telescope Targets and Observations</head><p>In order to quantify the GC populations of field UDGs we proposed for a Cycle 29 HST snapshot imaging program with WFC3. All the UDGs within 40 Mpc from  <ref type="bibr">van Dokkum et al. (2015)</ref> UDGs in Coma (red crosses) and LSB galaxies in Fornax (gray circles, <ref type="bibr">Venhola et al. 2017)</ref>. The horizontal dashed line indicates the standard r eff = 1.5 kpc cutoff for UDGs. The vertical dashed line at &#956; g,0 = 24 corresponds to the UDG surface brightness criterion of <ref type="bibr">van Dokkum et al. (2015)</ref>, while the more relaxed limit of van der <ref type="bibr">Burg et al. (2016)</ref> is approximated by the dotted line at &#956; g,0 = 23.2 (for a typical color of gr = 0.3). Nine of our targets meet the more stringent criteria, while 11 meet the more relaxed criteria. The horizontal red and black arrows indicate how the surface brightness values of the <ref type="bibr">van Dokkum et al. (2015)</ref> UDGs and our sample would shift (on average, based on their mean gr colors) if the r-band central surface brightness was used instead of the g band. As our objects are bluer their surface brightness does not increase as much between the g and r bands. <ref type="bibr">Janowiecki et al. (2019)</ref>, a total of 21 objects, were submitted for snapshot imaging (HST-16758; PI: M. Jones). The limit of 40 Mpc was to ensure that even for the most distant targets the turnover in the globular cluster luminosity function (GCLF) could be reached at high signal-to-noise ratio (S/N &gt; 5) in two filters in a single orbit. A total of 15 of these targets were observed during Cycle 29. Each target was observed for a total of 1000 s in two exposures in the F555W filter and 750 s in two exposures in F814W. Unfortunately, the observations of AGC 242019 lost tracking during the F555W exposures and the images were unusable, resulting in a sample of 14 targets (Table <ref type="table">1</ref>). The false color images from the two filters combined are shown for each target in Figure <ref type="figure">2</ref>.<ref type="foot">foot_1</ref> </p></div>
<div xmlns="http://www.tei-c.org/ns/1.0"><head n="2.2.">Revision of Photometry with DECaLS</head><p>The optical photometry of the parent H I-bearing UDGs sample was measured from SDSS images <ref type="bibr">(Leisman et al. 2017;</ref><ref type="bibr">Janowiecki et al. 2019</ref>) and size and surface brightness criteria were set to approximately match those of van der <ref type="bibr">Burg et al. (2016)</ref> for the full sample, and those of <ref type="bibr">van Dokkum et al. (2015)</ref> for a more restricted sample. Deeper Dark Energy Camera Legacy Survey (DECaLS; <ref type="bibr">Dey et al. 2019)</ref> images are now available for all our targets and we have therefore revised their photometry (Figure <ref type="figure">1</ref>).</p><p>The photometry for these UDGs is performed using AutoProf <ref type="bibr">(Stone et al. 2021)</ref>,aflexible, nonparametric fitting Python package that incorporates machine-learning methods (i.e., regularization) to improve upon previous fitting implementations. We retrieve 5&#8242; g-andr-band cutouts for each of the H I-bearing UDGs from DECaLS. Masks for these systems are generated using DeepScan <ref type="bibr">(Prole et al. 2018)</ref> and used in the surface brightness profile extraction. We take advantage of AutoProf&#700;s flexibility to measure the surface brightness profiles of our UDGs within circular apertures in a similar manner to <ref type="bibr">Leisman et al. (2017)</ref> and <ref type="bibr">Janowiecki et al. (2019)</ref>.</p><p>We then fit the extracted surface brightness profiles with an exponential function to estimate their central surface brightness and effective radii (Table <ref type="table">1</ref>) with uncertainties on these quantities estimated via bootstrap resampling.</p><p>With these deeper data, the uncertainties on the photometry have been significantly reduced and most objects have moved to slightly lower (brighter) &#956; g,0 values (central g-band surface brightness). Nine of the 14 targets meet the <ref type="bibr">van Dokkum et al. (2015)</ref> criteria for UDGs, while 11 meet the van der Burg et al.</p><p>(2016) criteria, and the remaining three are LSB galaxies near the border of these definitions (Figure <ref type="figure">1</ref>). Although the <ref type="bibr">van Dokkum et al. (2015)</ref> definition is more widely used, the van der <ref type="bibr">Burg et al. (2016)</ref> definition may be more appropriate for blue UDGs. First, they are blue because of recent star formation events, which will brighten their magnitude more in the g band than the r band. Thus, the r-band surface brightness is more representative of the total stellar content (see <ref type="bibr">Li et al. 2022)</ref>. Second, using central surface brightness makes most sense for galaxies with smooth light distributions, which can be accurately modeled with S&#233;rsic profiles. Most gas-rich, field UDGs have decidedly irregular and clumpy morphologies that only loosely follow an exponential profile. However, on average they do fall into the same extreme LSB regime as redder, smoother UDGs.</p><p>As the exact thresholds of surface brightness and size used to demarcate UDGs versus LSB galaxies is largely arbitrary, we will consider this small sample as a whole for the remainder of this work. However, we note that all of the qualitative findings would be unchanged if the sample were to be restricted to the nine objects that meet the most stringent criteria.</p><p>We also used the HST images to verify our new photometric measurements from DECaLS. We model each target galaxy using GALFIT <ref type="bibr">(Peng et al. 2002)</ref>, following the procedure from <ref type="bibr">Bennet et al. (2017)</ref>. These photometric fits were consistent with those from DECaLS, but generally found smaller radii, fainter integrated magnitudes, and significantly higher uncertainties. Thus, we elect to rely on the DECaLS results for the remainder of this work.  </p></div>
<div xmlns="http://www.tei-c.org/ns/1.0"><head n="3.">Globular Cluster Candidates</head><p>All individual WFC3 exposures (in both filters) were aligned and a combined source catalog was extracted for each target using DOLPHOT <ref type="bibr">(Dolphin 2000</ref><ref type="bibr">(Dolphin , 2016))</ref>. To select GCCs we begin by restricting the catalog to point-like sources (object type 1 and 2) with no photometry flags in either filter. Next a S/N minimum of 5 is enforced. Sources with more than 0.5 mag of additional flux (in the two filters combined) due to crowding are removed. The absolute sharpness value is required to be less than 0.25 in order to remove highly extended sources. Finally, a very loose roundness threshold of &lt;3 is enforced, which helps to remove any remaining diffraction spikes or other highly elongated features. The F555W and F814W magnitude of the remaining sources are corrected for Galactic extinction following Schlafly&amp; Finkbeiner (2011). The values of E(B -V ) are all less than 0.2, and most are less than 0.05. The corrected magnitudes are then converted to V and I (Vega) magnitudes following <ref type="bibr">Harris (2018)</ref>.</p><p>This results in a catalog of high-S/N, point-like objects that are concentrated around our target galaxy in each HST image. However, many of these sources correspond to young, starforming regions, not old GCs. Confusion between young star clusters and GCs is a well-known issue for the identification of GCs around late-type and irregular dwarfs. We adopt a simple color cut of V -I &gt; 0.85 to remove young clusters. <ref type="bibr">Seth et al. (2004)</ref> argue that any star cluster with a single stellar population that is older than 1 Gyr will have a color V -I &gt; 0.85, regardless of metallicity. We independently verified this cut using the PAdova and TRieste Stellar Evolution Code <ref type="bibr">(Bressan et al. 2012)</ref>. We also set an upper limit on the color of V -I &lt; 1.5, as almost no GCs are redder than this <ref type="bibr">(Brodie &amp; Strader 2006)</ref>.</p><p>To prevent contamination from bright stars we elect to only search for GCCs that are brighter than the turnover of the GCLF, M I,Vega = -8.12 (Miller &amp; Lotz 2007). We therefore enforce a magnitude range -8.12 &lt; M I,Vega &lt; -11.5, where the bright end roughly corresponds to the brightest known GCs. With this cut the number of GCs identified can simply be doubled to account for the uncounted fainter half of the GCLF.</p><p>Finally, to prevent the inclusion of any remaining background galaxies we use the Python package photutils to measure the concentration index of each source (in F814W) with apertures 4 and 8 pixels in diameter (as in <ref type="bibr">Beasley &amp; Trujillo 2016;</ref><ref type="bibr">Jones et al. 2021)</ref>. The restrictions on this concentration index are designed to be as relaxed as possible without including large numbers of background galaxies. The locus of point sources appears at approximately C 4-8 = 0.45 and we set the range as 0.2 &lt; C 4-8 &lt; 0.8. To verify that this criterion will not remove real GCs we modeled the expected concentration for a maximally extended GC. The largest GCs fit in <ref type="bibr">Larsen et al. (2001)</ref> have King profile core radii of &#8764;2.5 pc. We constructed a mock GC from a King profile with a core radius of 2.5 pc, and placed it at the nearest distance of any of the targets in our sample, 25 Mpc. This mock GC was then convolved with the average point-spread function (PSF) in the UVIS chip (using the WFC3/UVIS PSF models in F814W provided by the Space Telescope Science Institute), then the flux was extracted within the same-sized apertures as above. This gave a concentration index C 4-8 = 0.73, indicating that our concentration index cut should not exclude even the largest and nearest GCs. We also note that over all 14 targets only five potential GCCs are excluded solely on the basis of their concentration indices. We visually inspected all five and found that they were either diffraction spikes or clearly extended sources, likely background galaxies.</p><p>We inject artificial stars (point sources) into our images with true colors and magnitudes spread uniformly within our GCC selection box. These are successfully identified as GCCs approximately 90% of the time. The main reason for the eliminations is the large color uncertainties for the faintest objects, which can result in some objects falling outside the color range used for selection. In theory objects may also scatter into our selection box, however in practice most of the color-magnitude diagrams (CMDs; Appendix A) are so sparsely populated near the selection box that this is likely a negligible source on contaminants. We also see no change in the recovery rate toward the center of the target galaxies, where crowding might have been expected to cause issues. GCs (at least the brighter half of the GC population) appear to be sufficiently bright to prevent this crowding from playing a significant role.</p><p>Figure <ref type="figure">2</ref> shows the GCCs (small red circles) identified within twice the half-light radius (dashed green circles) of each target galaxy. As is immediately apparent upon inspection, there are very few GCs in these systems. <ref type="bibr">AGCs 189298, 191708, 258471</ref>, and 312297 all host a single GCC, while AGC 103435 has two (all meet the UDG criteria of <ref type="bibr">van Dokkum et al. 2015)</ref>. All the remaining nine systems have no GCCs.</p><p>In the case of AGC 191708 there are several potential GCCs near the edge of the area used to select objects that are likely associated with the galaxy (dashed green circle in Figure <ref type="figure">2</ref>).As all our targets are highly irregular a simple selection area of twice the half-light radius might not be suitable in some cases. However, in all other cases there are no other nearby sources meeting the GCC criteria, and so this does not pose a significant issue to GCC selection. Here we simply note that AGC 191708 might be slightly anomalous relative to the rest of the sample and could potentially host several GCs if its GC system was highly spatially extended.</p><p>We estimate the false-positive rate from field contaminants by counting the number of GCCs outside the encircled regions in the rest of the WFC3 field of view, and assuming that these are all false positives. After normalizing to the area of the search regions (dashed green circles, Figure <ref type="figure">2</ref>) we find that the targets with the highest false-positive rate are AGC 201993 and AGC 191708, with 0.8 and 0.9 false GCCs expected. However, no GCCs were identified in the former and only one in the latter. In all other cases the expectation is less than 0.25 false GCCs per target. Given these extremely low false-positive rates, we elect to make no correction to the GCC counts.</p><p>It is also possible that we have eliminated some real GCs with our color criterion (0.85 &lt; V -I &lt; 1.5). GCs typically follow a bimodal color distribution (e.g., <ref type="bibr">Brodie &amp; Strader 2006)</ref>, however only a few percent have colors V -I &lt; 0.85. The CMDs in the vicinity of each target galaxy also indicate that only AGC 312297 has a significant number of objects that are just blueward of our selection box (Appendix A), and these are likely young star clusters in the host galaxy rather than genuine GCs. Therefore, we also decide to neglect this systematic correction.</p><p>In Figure <ref type="figure">3</ref> we plot the number of GCs as a function of Vband absolute magnitude for our target sample compared to a broad sample of dwarf galaxies <ref type="bibr">(Harris et al. 2013)</ref>. We double the counts for each of our targets (with N GC &gt; 0) to approximately correct for the missing half of the GCLF that is fainter than our magnitude selection range. However, given the small number of GCs detected this is likely to overestimate the true number of GCs for any individual object, and we have thus plotted all values as upper limits. In the cases where no GCs were identified we set the upper limit estimate at N GC = 1. We have also plotted the mean value for the entire sample (with the errors showing the standard deviation), which falls just below zero on the plot as the mean number of GCs per galaxy is only 0.85 after applying the factor of 2 correction.</p><p>Our UDG and LSB dwarfs sample appears broadly consistent with the GC counts of other dwarf galaxies, but are toward the lower limit of the luminosity range sampled by <ref type="bibr">Harris et al. (2013)</ref>. We note that galaxies with N GC = 0 in the <ref type="bibr">Harris et al. (2013)</ref> sample are missing from this plot and it should not be used to compare to the objects in our sample where no GCs were detected. We consider these cases further in Section 4.</p><p>Coma Cluster UDGs are plotted with pink crosses. Although there is considerable scatter, on average these fall well above the N GC values of the dwarf galaxies in the comparison sample.</p><p>At the faintest magnitudes (M V &#8764;-12) there may be less difference between the Coma UDGs and normal dwarf galaxies, however the lack of objects near this magnitude in the <ref type="bibr">Harris et al. (2013)</ref> sample prevent a detailed comparison. In the magnitude range where most of our targets fall (-16 &lt; M V &lt; -14) the distribution of N GC for the Coma UDGs is clearly distinct from both normal dwarfs and our gasrich, field UDGs. However, it should be noted that the UDGs in our sample have markedly different colors from most cluster UDGs. If plotted in terms of stellar mass, then our UDGs would shift to the left and would overlap with some of the faintest cluster UDGs.</p></div>
<div xmlns="http://www.tei-c.org/ns/1.0"><head n="4.">Discussion</head><p>The results presented in the previous section indicate that gas-rich, field UDGs host relatively few GCs, in contrast to the many GC-rich UDGs typically found in cluster environments (e.g., <ref type="bibr">van Dokkum et al. 2017;</ref><ref type="bibr">Forbes et al. 2020;</ref><ref type="bibr">Lim et al. 2020)</ref>. In this section we discuss how these findings compare to normal dwarf galaxies and what implications this has for understanding the formation of field UDGs and their relation to UDGs in denser environments.</p></div>
<div xmlns="http://www.tei-c.org/ns/1.0"><head n="4.1.">Comparison to Normal Dwarf Galaxies</head><p>Using logistic regression, <ref type="bibr">Eadie et al. (2022)</ref> found that the "hurdle" stellar mass above which a galaxy is more likely to host at least GC than to host none is M * = 10 6.8 M e . Based on our stellar mass estimates of our targets (Table <ref type="table">1</ref> and Appendix B), all are above this threshold. From their regression model we would expect &#8764;75% of our targets to host at least one GC. We expect to miss about half of these as our GC selection method is only sensitive to the brighter half of the GCLF. Thus, our finding that five out of 14 targets have at least one GC is broadly in line with general expectations for dwarf galaxies in this stellar mass range, while those with a nonzero number of GCs identified all fall comfortably within the scatter of the <ref type="bibr">Harris et al. (2013)</ref> sample of normal galaxies shown in Figure <ref type="figure">3</ref>. Therefore, in terms of their GC populations, our target sample appears to be consistent with normal dwarf galaxies.</p><p>These UDGs are also consistent with normal field dwarfs in other respects. Using a sample of 12 galaxies drawn from the same parent population, <ref type="bibr">Gault et al. (2021)</ref> found that these UDGs follow the standard H I size-mass relations (e.g., <ref type="bibr">Wang et al. 2016)</ref>. <ref type="bibr">Janowiecki et al. (2019)</ref> also find that this population resides in an equivalent environment to other H Ibearing galaxies of a similar mass. These results suggest that whatever the root cause of their diffuse stellar distributions, it must be an internal process rather than one governed by environment and external effects, and it likely does not strongly influence the present-day distribution of the H I gas.</p><p>Together all these findings point to this subclass of UDGs being the extreme of the normal population of field dwarfs. Indeed, with the revised photometry using DECaLS imaging (Section 2.2 and Figure <ref type="figure">1</ref>), there appears to be a continuous distribution in surface brightness effective radius from the classical dwarf regime to the most extreme objects in the sample. We also note that UDGs and LSB galaxies in the Fornax cluster show a similar trend <ref type="bibr">(Venhola et al. 2017)</ref>.</p><p>However, there are some ways, aside from their diffuse stellar distributions, that these UDGs appear to differ from normal field galaxies. It has been pointed out by multiple works <ref type="bibr">(Leisman et al. 2017;</ref><ref type="bibr">Jones et al. 2018;</ref><ref type="bibr">Mancera Pi&#241;a et al. 2019</ref><ref type="bibr">, 2022)</ref> that they appear to be rotating more slowly than other gas-rich, field galaxies, and perhaps are even DM deficient. However, there are many uncertainties and biases that could potentially impact these findings (see <ref type="bibr">He et al. 2019)</ref>, and further investigation is required. <ref type="bibr">Kado-Fong et al. (2022a</ref><ref type="bibr">, 2022b)</ref> also recently suggested that these UDGs may have especially low star formation efficiencies (SFEs), relative to other field dwarfs, contributing to their ultra-diffuse appearance. However, we note that their control sample (which was optically selected rather than H I selected) was considerably less gas-rich than the UDG sample, which itself is fairly typical of other H I-selected galaxies in ALFALFA of similar stellar mass (see <ref type="bibr">Durbala et al. 2020)</ref>. This calls the result into question and again suggests that further work is needed to robustly contrast the star formation properties of field UDGs to normal field dwarfs.</p><p>In summary, in terms of their GC populations, environment, and H I sizes, gas-rich UDGs in the field are equivalent to other gas-rich dwarf galaxies, suggesting that they represent the extreme of a continuous distribution of surface brightness for field dwarf galaxies. However, further investigation is needed, particularly of their internal kinematics and SFEs, in order to constrain the mechanism(s) causing their diffuse stellar distributions (discussed further in Section 4.3).</p></div>
<div xmlns="http://www.tei-c.org/ns/1.0"><head n="4.2.">Could Field Ultra-diffuse Galaxies Represent the Progenitors of those in Denser Environments?</head><p>As soon as a large number of UDGs were identified in lowdensity environments, it was hypothesized that they could be representative of field UDGs from an earlier epoch that could have been the progenitors of present-day UDGs in clusters and groups <ref type="bibr">(Leisman et al. 2017)</ref>. More recently, <ref type="bibr">Junais &amp; Boselli (2022)</ref> argued that some blue UDGs in the outskirts of the Virgo cluster are being ram-pressure stripped and transforming into the red UDGs typically found in cluster environments. Given that blue UDGs in the field seem to make up a significant fraction of all UDGs (e.g., <ref type="bibr">Jones et al. 2018;</ref><ref type="bibr">Prole et al. 2019)</ref>, it is worth addressing the question: Were the UDGs found in present-day clusters and groups once similar to the blue, gas-rich UDGs found in the field?</p><p>Old star clusters are well suited to testing such a hypothesis as GC formation is thought to occur during the initial star formation episodes when a galaxy first forms (e.g., <ref type="bibr">Hudson et al. 2014)</ref>, and are therefore intricately connected to the halo mass of a galaxy. Thus, if two galaxy populations have strongly differing GC populations then their host halos must also differ, and they are unlikely to be physically related. Our findings clearly point to there being at least two pathways for forming UDGs, as the dearth of GCs in our field sample is incompatible with the GC-rich UDGs typically found in clusters. However, we note that UDGs in the Hydra cluster <ref type="bibr">(Iodice et al. 2020;</ref><ref type="bibr">La Marca et al. 2022</ref>) appear to host far fewer GCs than those in Coma, and some of these UDGs might have had progenitors similar to the field UDGs presented in this work.</p><p>The finding that field UDGs have far fewer GCs than most cluster UDGs is in tension with the hypothesis of <ref type="bibr">Junais &amp; Boselli (2022)</ref>, that blue UDGs in the outskirts of Virgo were previously field UDGs and are now transitioning to become red cluster UDGs. However, it is still possible that ram-pressure stripping of field UDGs may explain a small fraction of cluster UDGs, for example those at the lowest masses, which are less GC-rich (Figure <ref type="figure">3</ref>). An accounting of the GC systems of the <ref type="bibr">Junais &amp; Boselli (2022)</ref> sample would help to resolve this ambiguity.</p><p>Although it is clear that gas-rich, field UDGs cannot be representative of the progenitors of most cluster UDGs, the situation with UDGs in galaxy groups is more uncertain. <ref type="bibr">Somalwar et al. (2020)</ref> measured the number of GCs in a small sample (nine) of UDGs in two galaxy groups. They found that the UDGs spanned a range of GC system richness, with two objects being significantly above the distribution for normal dwarf galaxies of similar luminosity (green error bars, Figure <ref type="figure">3</ref>), though the remaining seven were consistent with normal dwarfs. The GC populations of the group UDGs DF2 and DF4 (van <ref type="bibr">Dokkum et al. 2018</ref><ref type="bibr">Dokkum et al. , 2019))</ref>, as well as <ref type="bibr">MATLAS-2019</ref><ref type="bibr">(Muller et al. 2021)</ref>, also reside at the upper limit or above those of normal dwarf galaxies. Also in a group environment, <ref type="bibr">Jones et al. (2021)</ref> found that two UDGs with evidence for tidal interactions <ref type="bibr">(Bennet et al. 2018</ref>) had GC systems consistent with normal dwarfs (dark orange error bars, Figure <ref type="figure">3</ref>). These results, albeit based on small samples, point to groups being an intermediate environment for UDGs, not just in the normal sense of neighboring galaxy density but also in terms of formation pathways for diffuse galaxies. At least some UDGs in groups appear analogous to those typically found in clusters (in terms of their GC populations), while the remainder have more typical GC systems and are presumably hosted by lowermass halos. <ref type="bibr">Jones et al. (2021)</ref> also argued that the two group UDGs in their study were most likely formed when normal field dwarfs fell into groups and were tidally heated, resulting in a more diffuse structure <ref type="bibr">(Bennet et al. 2018;</ref><ref type="bibr">Carleton et al. 2019;</ref><ref type="bibr">Tremmel et al. 2020)</ref>. The caveat was that, at the time, the properties of the GC systems of field UDGs were unknown. Our current finding, that gas-rich, field UDGs have GC systems that are consistent with normal dwarf galaxies, raises the possibility that these two UDGs might have been ultra-diffuse prior to falling into a group, and that the stellar streams they are adjacent to might not be indicative of the root cause of their diffuse structure <ref type="bibr">(Bennet et al. 2018)</ref>. We are actively pursuing a larger sample of similar UDGs to attempt to disentangle these possibilities.</p><p>We can also consider the fate of these specific UDGs and LSB galaxies, rather than what progenitors from a past epoch they could represent. <ref type="bibr">Janowiecki et al. (2019)</ref> found that H I- rich UDGs reside in the same environment as typical H I-rich dwarf galaxies of similar mass. That is, they are mostly centrals in their own low-mass halos <ref type="bibr">(Guo et al. 2017)</ref>. None of the 14 galaxies in our sample were matched to a known galaxy group by <ref type="bibr">Jones et al. (2020)</ref>. A visual inspection of the location of these particular galaxies reveals that those in the ALFALFA "Spring" sky are generally a few tens of degrees away from the Virgo cluster and mostly in the vicinity of filametary structures that extend from the cluster to the east and west (also extending to higher velocities). Those in the "Fall" sky appear to be in the foreground of the Pisces-Perseus Supercluster (about half way to the main structure), again in the vicinity of large-scale structures that mark the edge of a major foreground void. Thus, for these specific galaxies they will not be accreted on to a cluster for many billions of years (if ever), and are likely to remain as lone objects or perhaps join small groups.</p></div>
<div xmlns="http://www.tei-c.org/ns/1.0"><head n="4.3.">Comparison to Simulations and Models of Ultra-diffuse Galaxy Formation</head><p>Of the many proposed UDG formation mechanisms outlined in Section 1 only those that are internal mechanisms can be invoked to explain the existence of large numbers of UDGs in the field. Currently the most favored of these are those relying on repeated episodes of star formation feedback to redistribute matter to larger radii (e.g., <ref type="bibr">Chan et al. 2018;</ref><ref type="bibr">Di Cintio et al. 2017)</ref> or halos in the high-angular-momentum tail of the naturally occurring halo spin distribution <ref type="bibr">(Amorisco &amp; Loeb 2016;</ref><ref type="bibr">Rong et al. 2017)</ref>, with the angular momentum preventing an efficient collapse into a normally proportioned galaxy. Either of these models appear to be viable options for gas-rich, field UDGs. <ref type="bibr">Trujillo-Gomez et al. (2022)</ref> and <ref type="bibr">Danieli et al. (2022)</ref> also suggest that the formation of GCs themselves might be responsible for feedback that causes the diffuse structure of UDGs. However, for these models to explain our gas-rich, field UDGs they would likely need to host more GCs than we have identified, unless the majority have been lost.</p><p>Attempts have been made to measure the gas kinematics of these UDGs <ref type="bibr">(Mancera Pi&#241;a et al. 2019</ref><ref type="bibr">, 2020</ref><ref type="bibr">, 2022)</ref> as a means to probe their specific angular momenta. However, the poor resolution of most of the currently available data is problematic for drawing robust conclusions. In addition, even with accurate modeling of the kinematics of the baryons, there is no guarantee that this maps directly to the global DM halo angular momentum, which is not observable.</p><p>In the case of the star formation feedback models, much higher angular resolution kinematic data would be needed to directly identify the cored DM profiles that these models predict. Another means to test this model would be to obtain high-temporal-resolution star formation histories. However, all the objects in this sample are too distant to do this with HST, but it may be possible for some objects with the JWST. Given these challenges, it is important to ask what constraints can be drawn from the finding that these galaxies host few GCs.</p><p>When discussing the GC populations of Coma Cluster UDGs, <ref type="bibr">Saifollahi et al. (2022)</ref> highlighted that their high values of N GC /M * is an argument against forming UDGs purely by redistributing stellar mass to large radii, regardless of the exact mechanism for doing so. The formation pathway must include a mechanism that either increases the number of GCs or suppresses the expected stellar mass, or both. In the case of gas-rich, field UDGs, where the GC counts are equivalent to normal dwarf galaxies, this argument works in reverse. The correct explanation for their diffuse structure likely involves an almost purely redistributive process that does not significantly impact either GC formation or the overall buildup of stellar mass.</p><p>Complicating this somewhat, <ref type="bibr">Gault et al. (2021)</ref> found that the H I gas is not in a more extended distribution than is typical (for the total H I mass), therefore the redistribution process must only significantly affect stars (and potentially DM), not gas. As H I is generally much more spatially extended than the stars in most gas-rich galaxies, this may, for example, be possible with a cored DM halo, if the core is sufficiently compact as to not strongly influence the distribution of gas in the galaxy outskirts.</p><p>Recent hydrodynamical simulation results from IllustrisTNG <ref type="bibr">(Nelson et al. 2019a</ref>) indicate that late-type LSB galaxies form in higher-mass halos than higher-surface-brightness galaxies of the same stellar mass <ref type="bibr">(Perez-Montano et al. 2022</ref>). Expanding on this, <ref type="bibr">Benavides et al. (2022)</ref> found that UDGs in the TNG50 simulation <ref type="bibr">(Nelson et al. 2019b;</ref><ref type="bibr">Pillepich et al. 2019</ref>) are also skewed toward higher halo masses than normal dwarf galaxies, regardless of environment. The stellar mass estimates for the UDGs in our sample are typically &#61541; &#187; MM log 7.5 * , which, for a field UDG, corresponds to a halo mass &#61541; &#187; MM log 10.5 200 , according to <ref type="bibr">Benavides et al. (2022)</ref>.<ref type="foot">foot_2</ref> Zaritsky (2022) measured the linear relationship between the number of GCs and total galaxy mass, finding that on average there is 1 GC per (2.9 &#177; 0.3) &#215; 10 9 M e of total galaxy mass. If we use the halo mass estimate above as the total mass for our UDGs, then we would expect the UDGs in our sample to typically host 10.9 &#177; 1.1 GCs (note that this ignores the uncertainty in the halo mass estimate). Thus, the relative lack of GCs that we find is in clear tension with the expectation from TNG50 <ref type="bibr">(Benavides et al. 2022)</ref>. Given that the GC counts that we find appear to be compatible with normal dwarf galaxies, this suggests that these UDGs are in fact not hosted in DM halos that are more massive than those of other galaxies of similar stellar mass. If there are many unidentified field UDGs (analogous to DGSAT I; Section 4.4), then the findings of <ref type="bibr">Benavides et al. (2022)</ref> may offer an explanation, but this is not consistent with gas-rich, field UDGs. <ref type="bibr">Wright et al. (2021)</ref> reported an alternative formation pathway for field UDGs in the Romulus25 simulation <ref type="bibr">(Tremmel et al. 2017)</ref>. In this case UDGs were formed primarily through low-mass galaxy mergers that redistributed star formation more toward galaxy outskirts. This produced simulated field UDGs that have typical star formation rates and colors for field galaxies of similar stellar masses. The abundance of field UDGs in this model also matches quite well with estimates from <ref type="bibr">Jones et al. (2018)</ref>. As the relation between halo mass and N GC is linear <ref type="bibr">(Zaritsky 2022)</ref>, this scenario should also result in UDGs with GC systems comparable to normal dwarfs of similar masses.</p></div>
<div xmlns="http://www.tei-c.org/ns/1.0"><head n="4.4.">Comparison to DGSAT I</head><p>The UDG DGSAT I (Martinez-Delgado et al. 2016) is one of the few known quenched field UDGs. It is located in the vicinity of the Pisces-Perseus Supercluster at a distance of approximately 80 Mpc. A recent study of its GC system <ref type="bibr">(Janssens et al. 2022)</ref> produced the opposite finding to our sample of gas-rich, field UDGs: DGSAT I hosts a rich (N GC = 12 &#177; 2) and compact GC system, much more in line with UDGs in clusters.</p><p>It is possible that DGSAT I-like objects are the progenitors of cluster UDGs; however, it is a peculiar object even among UDGs, and such a conclusion would be premature without a larger sample of similar quenched field UDGs being identified first. What is clear is that DGSAT I did not form via the same pathway as the gas-rich UDGs we consider in this work. As discussed by <ref type="bibr">Janssens et al. (2022)</ref>, the most straightforward explanation may be that DGSAT I is a backsplash object that formed via the same mechanism as cluster UDGs.</p></div>
<div xmlns="http://www.tei-c.org/ns/1.0"><head n="5.">Conclusions</head><p>We have imaged 14 gas-rich, field UDGs and LSB galaxies with HST WFC3, and selected GCCs based on color, magnitude, and concentration index. We find strikingly few candidates, in stark contrast to the GC-rich UDGs typically found in galaxy clusters. Nine of our 14 targets have no GCCs brighter than the turnover magnitude of the dwarf galaxy GCLF (M I,Vega = -8.12).</p><p>These low GC counts are consistent with expectations for normal dwarf galaxies in a similar stellar mass range ( &#61541; MM log 7.5 *</p><p>), and suggest that the formation process driving the diffuse structure of these galaxies is primarily a redistributive process that moves stars to larger radii, without significantly impacting the formation of GCs, the long-term buildup of stellar mass, or the present-day distribution of neutral gas.</p><p>Given the small number of GCs in these field UDGs and LSB galaxies, they cannot represent the progenitors of red, GCrich UDGs in clusters, which presumably formed in highermass halos. However, they may be the progenitors of some UDGs in group environments which seem to exhibit a broad range of GC richness.</p><p>We thank the anonymous referee for their comments, which helped to improve this letter. This </p></div>
<div xmlns="http://www.tei-c.org/ns/1.0"><head>Appendix A Color-Magnitude Diagrams</head><p>The CMDs of point-like objects within twice the effective radius of each target galaxy are shown in Figure <ref type="figure">4</ref>.</p><p>In most cases there are very few sources in the expected magnitude range of GCs, and neither the color nor concentration index criteria play a significant role in selecting GCCs. Three exceptions appear to be AGCs 181474, 312297, and 749387. For AGC 181474 there is a cluster of sources (0 &lt; V -I &lt; 0.75 and 23 &lt; I &lt; 25) in the CMD that are just blueward of the selection box and fail the concentration index criteria. However, these are almost exclusively artifacts in the diffraction spikes of the two bright stars (Figure <ref type="figure">2</ref>) near the target galaxy. The same is true for AGC 749387, except the location of the artifacts in CMD is different (1 &lt; V -I &lt; 2 and 21.5 &lt; I &lt; 22.5). In the case of AGC 312297 there are a number of sources in the magnitude range expected for GCs Points (red or black) indicate sources that meet the concentration index criterion and black crosses are those that fail it. The gray rectangular outline shows the colormagnitude parameter space used to select GCCs, and red points are those sources that meet all the criteria. but just blueward of the selection box. These are likely young star clusters and H II regions, not old GCs.</p></div><note xmlns="http://www.tei-c.org/ns/1.0" place="foot" xml:id="foot_0"><p>The Astrophysical Journal Letters, 942:L5 (13pp), 2023 January 1 Jones et al.</p></note>
			<note xmlns="http://www.tei-c.org/ns/1.0" place="foot" n="11" xml:id="foot_1"><p>These images are available in the Mikulski Archive for Space Telescopes, 10.17909/6n6q-ke17.</p></note>
			<note xmlns="http://www.tei-c.org/ns/1.0" place="foot" n="12" xml:id="foot_2"><p>We note that this mass is at the lower extreme of their UDGs in their sample.</p></note>
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