The challenges associated with nowcasting quasi-linear convective system (QLCS) tornadoes are well documented. One key challenge is that QLCS tornadoes typically develop within mesovortices (MVs), but not all MVs are tornadic. This study used radar and in situ Pod data collected during the Propagation, Evolution, and Rotation in Linear Storms (PERiLS) field campaign to examine the characteristics that differentiate tornadic (TOR), wind-damaging (WD), and nondamaging (ND) MVs at various stages in their lifetimes and to investigate the low-level structure of QLCS MVs. Thirty-one QLCS MVs were manually identified and cataloged using the lowest elevation scans of the nearest WSR-88D and C-band on Wheels (COW) radars during the two years of PERiLS. TOR MVs, over their entire lifetimes, had stronger rotational velocities (Vrots), smaller diameters, and slightly longer lifetimes compared to WD and ND MVs. When MVs were analyzed during their pretornadic, predamaging, and prewarning phases (prephases), TOR and WD MVs had similar Vrots; however, TOR MVs typically had smaller diameters and contracted leading up to tornadogenesis, which could benefit nowcasters. In five cases, MVs were observed at the lowest WSR-88D elevation scans but were not visible in the COW data; the MV structure at different elevation angles for one case is presented. Eight Pods showed evidence of MV intercepts, demonstrated most notably by decreases in pressure. COW data, along with relatively weak wind speeds measured by Pods that collected data on MVs, suggest that vertical variations in low-level MV structure and strength can exist, which may not be adequately captured by the WSR-88D network.