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The latitudinal temperature gradient is a fundamental state parameter of the climate system tied to the dynamics of heat transport and radiative transfer. Thus, it is a primary target for temperature proxy reconstructions and global climate models. However, reconstructing the latitudinal temperature gradient in past climates remains challenging due to the scarcity of appropriate proxy records and large proxy–model disagreements. Here, we develop methods leveraging an extensive compilation of planktonic foraminifera δ 18 O to reconstruct a continuous record of the latitudinal sea-surface temperature (SST) gradient over the last 95 million years (My). We find that latitudinal SST gradients ranged from 26.5 to 15.3 °C over a mean global SST range of 15.3 to 32.5 °C, with the highest gradients during the coldest intervals of time. From this relationship, we calculate a polar amplification factor (PAF; the ratio of change in >60° S SST to change in global mean SST) of 1.44 ± 0.15. Our results are closer to model predictions than previous proxy-based estimates, primarily because δ 18 O-based high-latitude SST estimates more closely track benthic temperatures, yielding higher gradients. The consistent covariance of δ 18 O values in low- and high-latitude planktonic foraminifera and in benthic foraminifera, across numerous climate states, suggests a fundamental constraint on multiple aspects of the climate system, linking deep-sea temperatures, the latitudinal SST gradient, and global mean SSTs across large changes in atmospheric CO 2 , continental configuration, oceanic gateways, and the extent of continental ice sheets. This implies an important underlying, internally driven predictability of the climate system in vastly different background states. 

The widening of the South Atlantic Basin led to the reorganization of regional atmospheric and oceanic circulations. However, the response of the Atlantic Intertropical Convergence Zone (ITCZ), and South American and African monsoons across paleoclimate states, especially under constant paleogeographic and climatic changes, has not been well understood. Here we report on paleoclimate simulations of the Cenomanian (∼95 Ma), early Eocene (∼55 Ma), and middle Miocene (∼14 Ma) using the Community Earth System Model version 1.2 to understand how the migration of the South American and African continents to their modern‐day positions, uplift of the Andes and East African Rift Zone, and the decline of atmospheric CO₂changed the Atlantic ITCZ, and the South American and African monsoons and rainforests. Our work demonstrates that the South Atlantic widening developed the Atlantic ITCZ. The South Atlantic widening and Andean orogeny led to a stronger South American monsoon. We find the orogeny of the East African Rift Zone is the primary mechanism that strengthened the East African monsoon, whereas the West African monsoon became weaker through time as West Africa migrated toward the subtropics and CO₂levels fell below 500 ppm. We utilize the Köppen‐Geiger Climate Classification as an indicator for maximum rainforest extent. We find that during the Cenomanian and early Eocene, a Pan‐African rainforest existed, while the Amazon rainforest was restricted toward the northwestern corner of South America. During the middle Miocene, the Pan‐African rainforest was reduced to near its modern‐day size, while the Amazon rainforest expanded eastward.