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Abstract Physics‐based atmosphere‐land models with prescribed sea surface temperature have notable successes but also biases in their ability to represent atmospheric variability compared to observations. Recently, AI emulators and hybrid models have emerged with the potential to overcome these biases, but still require systematic evaluation against metrics grounded in fundamental atmospheric dynamics. We evaluate the representation of four atmospheric variability benchmarking metrics in a fully data‐driven AI emulator (ACE2‐ERA5) and hybrid model (NeuralGCM). The hybrid model and emulator can capture the spectra of large‐scale tropical waves and extratropical eddy‐mean flow interactions, including critical levels. However, both struggle to capture the timescales associated with quasi‐biennial oscillation (QBO, ∼28 months) and Southern annular mode propagation (∼150 days). These dynamical metrics serve as an initial benchmarking tool to inform AI model development and understand their limitations, which may be essential for out‐of‐distribution applications (e.g., extrapolating to unseen climates). 

Advances in quantitative genetics have enabled researchers to identify genomic regions associated with changes in phenotype. However, genomic regions can contain hundreds to thousands of genes, and progressing from genomic regions to candidate genes is still challenging. In genome-wide association studies (GWAS) measuring elemental accumulation (ionomic) traits, a mere 5% of loci are associated with a known ionomic gene - indicating that many causal genes are still unknown. To select candidates for the remaining 95% of loci, we developed a method to identify conserved genes underlying GWAS loci in multiple species. For 19 ionomic traits, we identified 14,336 candidates across Arabidopsis, soybean, rice, maize, and sorghum. We calculated the likelihood of candidates with random permutations of the data and determined that most of the top 10% of candidates were orthologous genes linked to GWAS loci across all five species. The candidate list also includes orthologous genes with previously established ionomic functions in Arabidopsis and rice. Our methods highlight the conserved nature of ionomic genetic regulators and enable the identification of previously unknown ionomic genes.