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Seismic anisotropy produced by aligned olivine in oceanic lithosphere offers a window into mid‐ocean ridge (MOR) dynamics. Yet, interpreting anisotropy in the context of grain‐scale deformation processes and strain observed in laboratory experiments and natural olivine samples has proven challenging due to incomplete seismological constraints and length scale differences spanning orders of magnitude. To bridge this observational gap, we estimate an in situ elastic tensor for oceanic lithosphere using co‐located compressional‐ and shear‐wavespeed anisotropy observations at the NoMelt experiment located on ∼70 Ma seafloor. The elastic model for the upper 7 km of the mantle, NoMelt_SPani7, is characterized by a fast azimuth parallel to the fossil‐spreading direction, consistent with corner‐flow deformation fabric. We compare this model with a database of 123 petrofabrics from the literature to infer olivine crystallographic orientations and shear strain accumulated within the lithosphere. Direct comparison to olivine deformation experiments indicates strain accumulation of 250%–400% in the shallow mantle. We find evidence for D‐type olivine lattice‐preferred orientation (LPO) with fast [100] parallel to the shear direction and girdled [010] and [001] crystallographic axes perpendicular to shear. D‐type LPO implies similar amounts of slip on the (010)[100] and (001)[100] easy slip systems during MOR spreading; we hypothesize that grain‐boundary sliding during dislocation creep relaxes strain compatibility, allowing D‐type LPO to develop in the shallow lithosphere. Deformation dominated by dislocation‐accommodated grain‐boundary sliding (disGBS) has implications for in situ stress and grain size during MOR spreading and implies grain‐size dependent deformation, in contrast to pure dislocation creep.

The relative motion of the lithosphere with respect to the asthenosphere implies the existence of a boundary zone that accommodates shear between the rigid plates and flowing mantle. This shear zone is typically referred to as the lithosphere‐asthenosphere boundary (LAB). The width of this zone and the mechanisms accommodating shear across it have important implications for coupling between mantle convection and surface plate motion. Seismic observations have provided evidence for several physical mechanisms that might help enable relative plate motion, but how these mechanisms each contribute to the overall accommodation of shear remains unclear. Here we present receiver function constraints on the discontinuity structure of the oceanic upper mantle at the NoMelt site in the central Pacific, where local constraints on shear velocity, anisotropy, conductivity, and attenuation down to ∼300 km depth provide a comprehensive picture of upper mantle structure. We image a seismic discontinuity with a Vsv decrease of 4.5% or more over a 0–20 km thick gradient layer centered at a depth of ∼65 km. We associate this feature with the Gutenberg discontinuity (G), and interpret our observation of G as resulting from strain localization across a dehydration boundary based on the good agreement between the discontinuity depth and that of the dry solidus. Transitions in Vsv, azimuthal anisotropy, conductivity, and attenuation observed at roughly similar depths suggest that the G discontinuity represents a region of localized strain within a broader zone accommodating shear between the lithosphere and asthenosphere.

A magnetotelluric survey in the Barotse Basin of western Zambia shows clear evidence for thinned lithosphere beneath an orogenic belt. The uppermost asthenosphere, at a depth of 60–70 km, is highly conductive, suggestive of the presence of a small amount of partial melt, despite the fact that there is no surface expression of volcanism in the region. Although the data support the presence of thicker cratonic lithosphere to the southeast of the basin, the lithospheric thickness is not well resolved and models show variations ranging from ~80 to 150 km in this region. Similarly variable is the conductivity of the mantle beneath the basin and immediately beneath the cratonic lithosphere to the southeast, although the conductivity is required to be elevated compared to normal lithospheric mantle. In a general sense, two classes of model are compatible with the magnetotelluric data: one with a moderately conductive mantle and one with more elevated conductivities. This latter class would be consistent with the impingement of a stringer of plume‐fed melt beneath the cratonic lithosphere, with the melt migrating upslope to thermally erode lithosphere beneath the orogenic belt that is overlain by the Barotse Basin. Such processes are potentially important for intraplate volcanism and also for development or propagation of rifting as lithosphere is thinned and weakened by melt. Both models show clear evidence for thinning of the lithosphere beneath the orogenic belt, consistent with elevated heat flow data in the region.