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Adaptive capacity can present challenges for modelling as it encompasses multiple ecological and evolutionary processes such as natural selection, genetic drift, gene flow and phenotypic plasticity. Spatially explicit, individual-based models provide an outlet for simulating these complex interacting eco-evolutionary processes. We expanded the existing Cost-Distance Meta-POPulation (CDMetaPOP) framework with inducible plasticity modelled as a habitat selection behaviour, using temperature or habitat quality variables, with a genetically based selection threshold conditioned on past individual experience. To demonstrate expected results in the new module, we simulated hypothetical populations and then evaluated model performance in populations of redband trout (Oncorhynchus mykiss gairdneri) across three watersheds where temperatures induce physiological stress in parts of the stream network. We ran simulations using projected warming stream temperature data under four scenarios for alleles that: (1) confer thermal tolerance, (2) bestow plastic habitat selection, (3) give both thermal tolerance and habitat selection preference and (4) do not provide either thermal tolerance or habitat selection. Inclusion of an adaptive allele decreased declines in population sizes, but this impact was greatly reduced in the relatively cool stream networks. As anticipated with the new module, high-temperature patches remained unoccupied by individuals with the allele operating plastically after exposure to warm temperatures. Using complete habitat avoidance above the stressful temperature threshold, habitat selection reduced the overall population size due to the opportunity cost of avoiding areas with increased, but not guaranteed, mortality. Inclusion of plasticity within CDMetaPOP will provide the potential for genetic or plastic traits and ‘rescue’ to affect eco-evolutionary dynamics for research questions and conservation applications. 

Abstract Patterns of energy and available moisture can vary over small (<1 km) distances in mountainous terrain. Information on fuel and soil moisture conditions that resolves this variation could help to inform fire and drought management decisions. Here, we describe the development of TOPOFIRE, a web-based mapping system designed to provide finely resolved information on soil water balance, drought, and wildfire danger information for the contiguous United States. We developed 8-arc-second-resolution (~250 meter) daily historical, near real-time, and 4-day forecast radiation, temperature, humidity, and snow water equivalent data and used these grids to calculate a suite of drought and wildfire danger indices. Large differences in shortwave radiation and surface air temperature with aspect contribute to greater snow accumulation and delays in melt timing on north-facing slopes, delaying fuel conditioning on shaded slopes. These datasets will help advance our understanding of the role of topography in wildland fire spread and ecological effects. Integration with national programs like the Wildland Fire Assessment System, the Wildland Fire Decision Support System, and drought early warning systems could support more proactive management of wildland fires and refine the characterization of drought in mountainous regions of the United States.