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Abstract Freshwater from the Arctic participates in the globally important Atlantic Meridional Overturning Circulation (AMOC). We use high‐resolution, in situ observations of dissolved organic matter (DOM) fluorescence to trace the origins of freshwater and organic carbon in the densest component of the AMOC, namely Denmark Strait Overflow Water (DSOW). We find a distinct terrestrial DOM signal in DSOW and trace it upstream to the Siberian shelves in the Arctic Ocean. This implies a riverine origin of freshwater in DSOW. We estimate that the Siberian Shelf water contribution constitutes approximately 1% of DSOW. Ocean circulation modeling confirms the inferred pathway and highlights Denmark Strait as an important location for the entrainment of the riverine signal into DSOW. Our proposed method can be deployed on a range of observing systems to elucidate freshwater dispersion across the Arctic and subarctic, thereby contributing to the broader discussion on freshwater impacts and organic carbon sequestration in the AMOC. 

American bison (Bison bison) were nearly hunted to extinction by the 20th century. This rapid decline and the fragmented nature of remnant populations pose challenges to their recovery and resilience, as does human-facilitated admixture between bison subspecies and cattle (Bos taurus). To contextualize current diversity, we sequenced 115 ancient and 45 modern bison genomes from the past ~20,000 years. Our results show that past bison populations were connected, in contrast to structured modern herds. Modern wood bison (B. b. athabascae), a northerly distributed subspecies, carry plains bison (B. b. bison) ancestry from 1920s translocations, and many sampled bison lack cattle ancestry. Our findings reveal the legacy of human impacts on bison and highlight the applicability of ancient DNA for guiding wildlife restoration. 

Growing season temperatures play a crucial role in controlling treeline elevation at regional to global scales. However, understanding of treeline dynamics in response to long-term changes in temperature is limited. In this study, we analyze pollen, plant macrofossils, and charcoal preserved in organic layers within a 10,400-year-old ice patch and in sediment from a 6000-year-old wetland located above present-day treeline in the Beartooth Mountains, Wyoming, to explore the relationship between Holocene climate variability and shifts in treeline elevation. Pollen data indicate a lower-than-present treeline between 9000 and 6200 cal yr BP during the warm, dry summer and cold winter conditions of the early Holocene. Increases in arboreal pollen at 6200 cal yr BP suggest an upslope treeline expansion when summers became cooler and wetter. A possible hiatus in the wetland record at ca. 4200–3000 cal yr BP suggests increased snow and ice cover at high elevations and a lowering of treeline. Treeline position continued to fluctuate with growing season warming and cooling during the late-Holocene. Periods of high fire activity correspond with times of increased woody cover at high elevations. The two records indicate that climate was an important driver of vegetation and treeline change during the Holocene. Early Holocene treeline was governed by moisture limitations, whereas late-Holocene treeline was sensitive to increases in growing season temperatures. Climate projections for the region suggest warmer temperatures could decrease effective growing season moisture at high elevations resulting in a reduction of treeline elevation. 

Arctic Ocean gateway fluxes play a crucial role in linking the Arctic with the global ocean and affecting climate and marine ecosystems. We reviewed past studies on Arctic–Subarctic ocean linkages and examined their changes and driving mechanisms. Our review highlights that radical changes occurred in the inflows and outflows of the Arctic Ocean during the 2010s. Specifically, the Pacific inflow temperature in the Bering Strait and Atlantic inflow temperature in the Fram Strait hit record highs, while the Pacific inflow salinity in the Bering Strait and Arctic outflow salinity in the Davis and Fram straits hit record lows. Both the ocean heat convergence from lower latitudes to the Arctic and the hydrological cycle connecting the Arctic with Subarctic seas were stronger in 2000–2020 than in 1980–2000. CMIP6 models project a continuing increase in poleward ocean heat convergence in the 21st century, mainly due to warming of inflow waters. They also predict an increase in freshwater input to the Arctic Ocean, with the largest increase in freshwater export expected to occur in the Fram Strait due to both increased ocean volume export and decreased salinity. Fram Strait sea ice volume export hit a record low in the 2010s and is projected to continue to decrease along with Arctic sea ice decline. We quantitatively attribute the variability of the volume, heat, and freshwater transports in the Arctic gateways to forcing within and outside the Arctic based on dedicated numerical simulations and emphasize the importance of both origins in driving the variability. 

Abstract. The Arctic Mediterranean (AM) is the collective name forthe Arctic Ocean, the Nordic Seas, and their adjacent shelf seas. Water enters into thisregion through the Bering Strait (Pacific inflow) and through the passages across theGreenland–Scotland Ridge (Atlantic inflow) and is modified within the AM. The modifiedwaters leave the AM in several flow branches which are grouped into two differentcategories: (1) overflow of dense water through the deep passages across theGreenland–Scotland Ridge, and (2) outflow of light water – here termed surface outflow– on both sides of Greenland. These exchanges transport heat and salt into and out ofthe AM and are important for conditions in the AM. They are also part of the global oceancirculation and climate system. Attempts to quantify the transports by various methodshave been made for many years, but only recently the observational coverage has becomesufficiently complete to allow an integrated assessment of the AM exchanges based solelyon observations. In this study, we focus on the transport of water and have collecteddata on volume transport for as many AM-exchange branches as possible between 1993 and2015. The total AM import (oceanic inflows plusfreshwater) is found to be 9.1 Sv (sverdrup,1 Sv =10⁶ m³ s⁻¹) with an estimated uncertainty of 0.7 Sv and hasthe amplitude of the seasonal variation close to 1 Sv and maximum import in October.Roughly one-third of the imported water leaves the AM as surface outflow with theremaining two-thirds leaving as overflow. The overflow water is mainly produced frommodified Atlantic inflow and around 70 % of the total Atlantic inflow is convertedinto overflow, indicating a strong coupling between these two exchanges. The surfaceoutflow is fed from the Pacific inflow and freshwater (runoff and precipitation), but isstill approximately two-thirds of modified Atlantic water. For the inflowbranches and the two main overflow branches (Denmark Strait and Faroe Bank Channel),systematic monitoring of volume transport has been established since the mid-1990s, andthis enables us to estimate trends for the AM exchanges as a whole. At the 95 %confidence level, only the inflow of Pacific water through the Bering Strait showed astatistically significant trend, which was positive. Both the total AM inflow and thecombined transport of the two main overflow branches also showed trends consistent withstrengthening, but they were not statistically significant. They do suggest, however,that any significant weakening of these flows during the last two decades is unlikely andthe overall message is that the AM exchanges remained remarkably stable in the periodfrom the mid-1990s to the mid-2010s. The overflows are the densest source water for thedeep limb of the North Atlantic part of the meridional overturning circulation (AMOC),and this conclusion argues that the reported weakening of the AMOC was not due tooverflow weakening or reduced overturning in the AM. Although the combined data set hasmade it possible to establish a consistent budget for the AM exchanges, the observationalcoverage for some of the branches is limited, which introduces considerable uncertainty.This lack of coverage is especially extreme for the surface outflow through the DenmarkStrait, the overflow across the Iceland–Faroe Ridge, and the inflow over the Scottishshelf. We recommend that more effort is put into observing these flows as well asmaintaining the monitoring systems established for the other exchange branches.