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Abstract Most great earthquakes on subduction zone plate boundaries have large coseismic slip concentrated along the contact between the subducting slab and the upper plate crust. On 4 March 2021, a magnitude 7.4 foreshock struck 1 hr 47 min before a magnitude 8.1 earthquake along the northern Kermadec island arc. The mainshock is the largest well‐documented underthrusting event along the ∼2,500‐km long Tonga‐Kermadec subduction zone. Using teleseismic, geodetic, and tsunami data, we find that all substantial coseismic slip in the mainshock is located along the mantle/slab interface at depths from 20 to 55 km, with the large foreshock nucleating near the down‐dip edge. Smaller foreshocks and most aftershocks are located up‐dip of the mainshock, where substantial prior moderate thrust earthquake activity had occurred. The upper plate crust is ∼17 km thick in northern Kermadec with only moderate‐size events along the crust/slab interface. A 1976 sequence withM_Wvalues of 7.9, 7.8, 7.3, 7.0, and 7.0 that spanned the 2021 rupture zone also involved deep megathrust rupture along the mantle/slab contact, but distinct waveforms exclude repeating ruptures. Variable waveforms for eight deep M6.9+ thrusting earthquakes since 1990 suggest discrete slip patches distributed throughout the region. The ∼300‐km long plate boundary in northern Kermadec is the only documented subduction zone region where the largest modeled interplate earthquakes have ruptured along the mantle/slab interface, suggesting that local frictional properties of the putatively hydrated mantle wedge may involve a dense distribution of Antigorite‐rich patches with high slip rate velocity weakening behavior in this locale.