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Extending the benefits of online reading to people with reading disabilities such as dyslexia requires broader research on reading behavior in addition to existing small-scale eye-tracking studies. We conduct the first large-scale mixed-methods study of the unique reading challenges of people with dyslexia. We combine in-person interviews (N=6), online surveys (N=566) and a novel browser-based tool able to measure detailed reading behavior remotely on a controlled set of five pages (N=477) or as a browser extension (N=89) collecting long-term reading behavior data on self-selected pages. We find a variety of text and page layout factors that pose challenges to readers with and without dyslexia, and identify in-browser reading behaviors associated with dyslexia. Findings point toward improvements to technologies for identifying struggling readers, and to ways to improve the layout and appearance of online articles to improve reading ease for people with and without dyslexia.