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The structure of the critical zone (CZ) is a product of feedbacks among hydrologic, climatic, biotic, and chemical processes. Past research within snow‐dominated systems has shown that aspect‐dependent solar radiation inputs can produce striking differences in vegetation composition, topography, and soil depth between opposing hillslopes. However, far fewer studies have evaluated the role of microclimates on CZ development within rain‐dominated systems, especially below the soil and into weathered bedrock. To address this need, we characterized the CZ of a north‐facing and south‐facing slope within a first‐order headwater catchment located in central coast California. We combined terrain analysis of vegetation distribution and topography with soil pit characterization, geophysical surveys and hydrologic measurements between slope‐aspects. We documented denser vegetation and higher shallow soil moisture on north facing slopes, which matched previously documented observations in snow‐dominated sites. However, average topographic gradients were 24° and saprolite thickness was approximately 6 m across both hillslopes, which did not match common observations from the literature that showed widespread asymmetry in snow‐dominated systems. These results suggest that dominant processes for CZ evolution are not necessarily transferable across regions. Thus, there is a continued need to expand CZ research, especially in rain‐dominated and water‐limited systems. Here, we present two non‐exclusive mechanistic hypotheses that may explain these unexpected similarities in slope and saprolite thickness between hillslopes with opposing aspects.

 

Building on the successful Argo network of seafaring temperature and salinity sensors, work is underway to deploy 1,000 floats equipped to study ocean biogeochemistry in greater detail than ever. 

The northeastern United States experienced extensive deforestation during the seventeenth through twentieth centuries primarily for European agriculture, which peaked in the mid-nineteenth century, and followed by widespread farmstead abandonment and reforestation. Analysis of airborne light detection and ranging (LiDAR) data has revealed thousands of historical land-use features with topographic signatures across the landscape under the region’s now-dense forest canopy. This study investigates two different types of features—stone walls and relict charcoal hearths—both of which are associated with widespread deforestation in the region. Our results demonstrate that LiDAR is an effective tool in reconstructing and quantifying the distribution and magnitude of historical forest cover using these relict land use features as a reliable proxy. Furthermore, these methods allow for direct quantification of cumulative land clearing over time in each town, in addition to the extent, intensity, and spatial distribution of cleared land and forest cover. 

Abstract The genus Anax is a group of cosmopolitan dragonflies noted for its conspicuous migratory behaviours and large size. Here we present the first dated, species-level, multigene, molecular phylogeny for the group to test generic and species-limits, as well as the evolution of migration and range size. Using five mitochondrial and nuclear gene regions (COI, COI/COII, CYTB/ND1, ITS1 and PRMT) from 20 species, we reconstructed a phylogeny of Anax using both a Bayesian and maximum likelihood approach. We found that Anax (including its hypothesized sister group Hemianax) forms a monophyletic group, and that 12 out of 20 species tested positive for monophyly were also monophyletic. The monophyly of several species of Anax is less clear. Migratory behaviour, which is known to occur in at least nine species, is recovered as the ancestral behaviour, which was lost and subsequently gained at least three times. Geographic range size seems to be tightly associated with migratory behaviour.