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The evening primrose family (Onagraceae) includes 664 species (803 taxa) with a center of diversity in the Americas, especially western North America. Ongoing research in Onagraceae includes exploring striking variation in floral morphology, scent composition, and breeding system, as well as the role of these traits in driving diversity among plants and their interacting pollinators and herbivores. However, these efforts are limited by the lack of a comprehensive, well-resolved phylogeny. Previous phylogenetic studies based on a few loci strongly support the monophyly of the family and the sister relationship of the two largest tribes but fail to resolve several key relationships.

We used a target enrichment approach to reconstruct the phylogeny of Onagraceae using 303 highly conserved, low-copy nuclear loci. We present a phylogeny for Onagraceae with 169 individuals representing 152 taxa sampled across the family, including extensive sampling within the largest tribe, Onagreae. Deep splits within the family are strongly supported, whereas relationships among closely related genera and species are characterized by extensive conflict among individual gene trees.

This phylogenetic resource will augment current research projects focused throughout the family in genomics, ecology, coevolutionary dynamics, biogeography, and the evolution of characters driving diversification in the family.

 

Numerous studies have shown reduced performance in plants that are surrounded by neighbours of the same species^1,2, a phenomenon known as conspecific negative density dependence (CNDD)³. A long-held ecological hypothesis posits that CNDD is more pronounced in tropical than in temperate forests^4,5, which increases community stabilization, species coexistence and the diversity of local tree species^6,7. Previous analyses supporting such a latitudinal gradient in CNDD^8,9have suffered from methodological limitations related to the use of static data^10–12. Here we present a comprehensive assessment of latitudinal CNDD patterns using dynamic mortality data to estimate species-site-specific CNDD across 23 sites. Averaged across species, we found that stabilizing CNDD was present at all except one site, but that average stabilizing CNDD was not stronger toward the tropics. However, in tropical tree communities, rare and intermediate abundant species experienced stronger stabilizing CNDD than did common species. This pattern was absent in temperate forests, which suggests that CNDD influences species abundances more strongly in tropical forests than it does in temperate ones¹³. We also found that interspecific variation in CNDD, which might attenuate its stabilizing effect on species diversity^14,15, was high but not significantly different across latitudes. Although the consequences of these patterns for latitudinal diversity gradients are difficult to evaluate, we speculate that a more effective regulation of population abundances could translate into greater stabilization of tropical tree communities and thus contribute to the high local diversity of tropical forests.

 

A new surface-rupture-length (SRL) relationship as a function of magnitude [Formula: see text], fault thickness, and fault dip angle is presented in this article. The objective of this study is to model the change in scaling between unbounded and width-limited ruptures. This is achieved through the use of seismological-theory-based relationships for the average displacement scaling and the aid of dynamic fault rupture simulations to constrain the rupture width scaling. The empirical data set used in the development of this relationship is composed of [Formula: see text] events ranging from [Formula: see text] 5 to [Formula: see text] and SRL 1.1 to 432 km. The dynamic rupture simulations data set includes [Formula: see text] events ranging from [Formula: see text] 4.9 to 8.2 and [Formula: see text] 1 to 655 km. For the average displacement [Formula: see text] scaling, three simple models and two composite models were evaluated. The simple average displacement models are a square root of the rupture area [Formula: see text], a down-dip width [Formula: see text], and a rupture length [Formula: see text] proportional model. The two composite models have a [Formula: see text] scaling for unbounded ruptures and transition to [Formula: see text] and [Formula: see text] scaling for width-limited events, respectively. The empirical data favors a [Formula: see text] scaling for the entire regime (unbounded and width-limited ruptures) followed by a [Formula: see text] scaling for unbounded that changes to [Formula: see text] scaling for width-limited ruptures. The selected models exhibit better predictive performance compared to linear [Formula: see text] type models, especially in the large magnitude range, which is dominated by width-limited events. A comparison with published SRL models shows consistent scaling for different fault types and tectonic environments.

 

We present near-infrared (NIR) ground-basedY,J,H, andKimaging obtained in the James Webb Space Telescope (JWST) North Ecliptic Pole Time Domain Field (NEP TDF) using the MMT-Magellan Infrared Imager and Spectrometer on the MMT. These new observations cover a field of approximately 230 arcmin²inY,H, andK,and 313 arcmin²inJ. Using Monte Carlo simulations, we estimate a 1σdepth relative to the background sky of (Y, J, H, K) = (23.80, 23.53, 23.13, 23.28) in AB magnitudes for point sources at a 95% completeness level. These observations are part of the ground-based effort to characterize this region of the sky, supplementing space-based data obtained with Chandra, NuSTAR, XMM, AstroSat, Hubble Space Telescope, and JWST. This paper describes the observations and reduction of the NIR imaging and combines these NIR data with archival imaging in the visible, obtained with the Subaru Hyper-Suprime-Cam, to produce a merged catalog of 57,501 sources. The new observations reported here, plus the corresponding multiwavelength catalog, will provide a baseline for time-domain studies of bright sources in the NEP TDF.

 

One mechanism proposed to explain high species diversity in tropical systems is strong negative conspecific density dependence (CDD), which reduces recruitment of juveniles in proximity to conspecific adult plants. Although evidence shows that plant-specific soil pathogens can drive negative CDD, trees also form key mutualisms with mycorrhizal fungi, which may counteract these effects. Across 43 large-scale forest plots worldwide, we tested whether ectomycorrhizal tree species exhibit weaker negative CDD than arbuscular mycorrhizal tree species. We further tested for conmycorrhizal density dependence (CMDD) to test for benefit from shared mutualists. We found that the strength of CDD varies systematically with mycorrhizal type, with ectomycorrhizal tree species exhibiting higher sapling densities with increasing adult densities than arbuscular mycorrhizal tree species. Moreover, we found evidence of positive CMDD for tree species of both mycorrhizal types. Collectively, these findings indicate that mycorrhizal interactions likely play a foundational role in global forest diversity patterns and structure.

 

We derive the spatial and wavelength behavior of dust attenuation in the multiple-armed spiral galaxy VV 191b using backlighting by the superimposed elliptical system VV 191a in a pair with an exceptionally favorable geometry for this measurement. Imaging using the James Webb Space Telescope and Hubble Space Telescope spans the wavelength range 0.3–4.5μm with high angular resolution, tracing the dust in detail from 0.6–1.5μm. Distinct dust lanes continue well beyond the bright spiral arms, and trace a complex web, with a very sharp radial cutoff near 1.7 Petrosian radii. We present attenuation profiles and coverage statistics in each band at radii 14–21 kpc. We derive the attenuation law with wavelength; the data both within and between the dust lanes clearly favor a stronger reddening behavior (R=A_V/E_B−V≈ 2.0 between 0.6 and 0.9μm, approaching unity by 1.5μm) than found for starbursts and star-forming regions of galaxies. Power-law extinction behavior ∝λ^−βgivesβ= 2.1 from 0.6–0.9μm.Rdecreases at increasing wavelengths (R≈ 1.1 between 0.9 and 1.5μm), whileβsteepens to 2.5. Mixing regions of different column density flattens the wavelength behavior, so these results suggest a different grain population than in our vicinity. The NIRCam images reveal a lens arc and counterimage from a background galaxy atz≈ 1, spanning 90° azimuthally at 2.″8 from the foreground elliptical-galaxy nucleus, and an additional weakly lensed galaxy. The lens model and imaging data give a mass/light ratioM/L_B= 7.6 in solar units within the Einstein radius 2.0 kpc.

 

The Time Domain Field (TDF) near the North Ecliptic Pole in JWST’s continuous-viewing zone will become a premier “blank field” for extragalactic science. JWST/NIRCam data in a 16 arcmin²portion of the TDF identify 4.4μm counterparts for 62 of 63 3 GHz sources withS(3 GHz) > 5μJy. The one unidentified radio source may be a lobe of a nearby Seyfert galaxy, or it may be an infrared-faint radio source. The bulk properties of the radio-host galaxies are consistent with those found by previous work: redshifts range from 0.14–4.4 with a median redshift of 1.33. The radio emission arises primarily from star formation in ∼2/3 of the sample and from an active galactic nucleus (AGN) in ∼1/3, but just over half the sample shows evidence for an AGN either in the spectral energy distribution or by radio excess. All but three counterparts are brighter than magnitude 23 AB at 4.4μm, and the exquisite resolution of JWST identifies correct counterparts for sources for which observations with lower angular resolution would misidentify a nearby bright source as the counterpart when the correct one is faint and red. Up to 11% of counterparts might have been unidentified or misidentified absent NIRCam observations.

 

Oenothera sect. Calylophus is a North American group of 13 recognized taxa in the evening primrose family (Onagraceae) with an evolutionary history that may include independent origins of bee pollination, edaphic endemism, and permanent translocation heterozygosity. Like other groups that radiated relatively recently and rapidly, taxon boundaries within Oenothera sect. Calylophus have remained challenging to circumscribe. In this study, we used target enrichment, flanking noncoding regions, gene tree/species tree methods, tests for gene flow modified for target-enrichment data, and morphometric analysis to reconstruct phylogenetic hypotheses, evaluate current taxon circumscriptions, and examine character evolution in Oenothera sect. Calylophus. Because sect. Calylophus comprises a clade with a relatively restricted geographic range, we were able to extensively sample across the range of geographic, edaphic, and morphological diversity in the group. We found that the combination of exons and flanking noncoding regions led to improved support for species relationships. We reconstructed potential hybrid origins of some accessions and note that if processes such as hybridization are not taken into account, the number of inferred evolutionary transitions may be artificially inflated. We recovered strong evidence for multiple evolutionary origins of bee pollination from ancestral hawkmoth pollination, edaphic specialization on gypsum, and permanent translocation heterozygosity. This study applies newly emerging techniques alongside dense infraspecific sampling and morphological analyses to effectively reconstruct the recalcitrant history of a rapid radiation. [Gypsum endemism; Oenothera sect. Calylophus; Onagraceae; phylogenomics; pollinator shift; recent radiation; target enrichment.]

 

Using the first epoch of four-band NIRCam observations obtained by the James Webb Space Telescope (JWST) Prime Extragalactic Areas for Reionization and Lensing Science Program in the Spitzer IRAC Dark Field, we search for F150W and F200W dropouts. In 14.2 arcmin², we have found eight F150W dropouts and eight F200W dropouts, all brighter than 27.5 mag (the brightest being ∼24 mag) in the band to the red side of the break. As they are detected in multiple bands, these must be real objects. Their nature, however, is unclear, and characterizing their properties is important for realizing the full potential of JWST. If the observed color decrements are due to the Lyman break, these objects should be atz≳ 11.7 andz≳ 15.4, respectively. The color diagnostics show that at least four F150W dropouts are far away from the usual contaminators encountered in dropout searches (red galaxies at much lower redshifts or brown dwarf stars). While the diagnostics of the F200W dropouts are less certain due to the limited number of passbands, at least one of them is likely not a known type of contaminant, and the rest are consistent with either high-redshift galaxies with evolved stellar populations or old galaxies atz≈ 3–8. If a significant fraction of our dropouts are indeed atz≳ 12, we have to face the severe problem of explaining their high luminosities and number densities. Spectroscopic identifications of such objects are urgently needed.

 

Abstract We report the discovery of a new “changing-look” active galactic nucleus (CLAGN) event, in the quasar SDSS J162829.17+432948.5 at z = 0.2603, identified through repeat spectroscopy from the fifth Sloan Digital Sky Survey (SDSS-V). Optical photometry taken during 2020–2021 shows a dramatic dimming of Δ g ≈ 1 mag, followed by a rapid recovery on a timescale of several months, with the ≲2 month period of rebrightening captured in new SDSS-V and Las Cumbres Observatory spectroscopy. This is one of the fastest CLAGN transitions observed to date. Archival observations suggest that the object experienced a much more gradual dimming over the period of 2011–2013. Our spectroscopy shows that the photometric changes were accompanied by dramatic variations in the quasar-like continuum and broad-line emission. The excellent agreement between the pre- and postdip photometric and spectroscopic appearances of the source, as well as the fact that the dimmest spectra can be reproduced by applying a single extinction law to the brighter spectral states, favor a variable line-of-sight obscuration as the driver of the observed transitions. Such an interpretation faces several theoretical challenges, and thus an alternative accretion-driven scenario cannot be excluded. The recent events observed in this quasar highlight the importance of spectroscopic monitoring of large active galactic nucleus samples on weeks-to-months timescales, which the SDSS-V is designed to achieve.