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Determining how the galactic environment, especially the high gas densities and complex dynamics in bar-fed galaxy centers, alters the star formation efficiency (SFE) of molecular gas is critical to understanding galaxy evolution. However, these same physical or dynamical effects also alter the emissivity properties of CO, leading to variations in the CO-to-H₂conversion factor (α_CO) that impact the assessment of the gas column densities and thus of the SFE. To address such issues, we investigate the dependence ofα_COon the local CO velocity dispersion at 150 pc scales using a new set of dust-basedα_COmeasurements and propose a newα_COprescription that accounts for CO emissivity variations across galaxies. Based on this prescription, we estimate the SFE in a sample of 65 galaxies from the PHANGS–Atacama Large Millimeter/submillimeter Array survey. We find increasing SFE toward high-surface-density regions like galaxy centers, while using a constant or metallicity-basedα_COresults in a more homogeneous SFE throughout the centers and disks. Our prescription further reveals a mean molecular gas depletion time of 700 Myr in the centers of barred galaxies, which is overall three to four times shorter than in nonbarred galaxy centers or the disks. Across the galaxy disks, the depletion time is consistently around 2–3 Gyr, regardless of the choice ofα_COprescription. All together, our results suggest that the high level of star formation activity in barred centers is not simply due to an increased amount of molecular gas, but also to an enhanced SFE compared to nonbarred centers or disk regions.

 

Abstract The CO-to-H 2 conversion factor ( α CO ) is central to measuring the amount and properties of molecular gas. It is known to vary with environmental conditions, and previous studies have revealed lower α CO in the centers of some barred galaxies on kiloparsec scales. To unveil the physical drivers of such variations, we obtained Atacama Large Millimeter/submillimeter Array bands (3), (6), and (7) observations toward the inner ∼2 kpc of NGC 3627 and NGC 4321 tracing 12 CO, 13 CO, and C 18 O lines on ∼100 pc scales. Our multiline modeling and Bayesian likelihood analysis of these data sets reveal variations of molecular gas density, temperature, optical depth, and velocity dispersion, which are among the key drivers of α CO . The central 300 pc nuclei in both galaxies show strong enhancement of temperature T k ≳ 100 K and density n H 2 > 10 3 cm −3 . Assuming a CO-to-H 2 abundance of 3 × 10 −4 , we derive 4–15 times lower α CO than the Galactic value across our maps, which agrees well with previous kiloparsec-scale measurements. Combining the results with our previous work on NGC 3351, we find a strong correlation of α CO with low- J 12 CO optical depths ( τ CO ), as well as an anticorrelation with T k . The τ CO correlation explains most of the α CO variation in the three galaxy centers, whereas changes in T k influence α CO to second order. Overall, the observed line width and 12 CO/ 13 CO 2–1 line ratio correlate with τ CO variation in these centers, and thus they are useful observational indicators for α CO variation. We also test current simulation-based α CO prescriptions and find a systematic overprediction, which likely originates from the mismatch of gas conditions between our data and the simulations. 

Galactic bars can drive cold gas inflows towards the centres of galaxies. The gas transport happens primarily through the so-called bar dust lanes, which connect the galactic disc at kpc scales to the nuclear rings at hundreds of pc scales much like two gigantic galactic rivers. Once in the ring, the gas can fuel star formation activity, galactic outflows, and central supermassive black holes. Measuring the mass inflow rates is therefore important to understanding the mass/energy budget and evolution of galactic nuclei. In this work, we use CO datacubes from the PHANGS-ALMA survey and a simple geometrical method to measure the bar-driven mass inflow rate on to the nuclear ring of the barred galaxy NGC 1097. The method assumes that the gas velocity in the bar lanes is parallel to the lanes in the frame co-rotating with the bar, and allows one to derive the inflow rates from sufficiently sensitive and resolved position–position–velocity diagrams if the bar pattern speed and galaxy orientations are known. We find an inflow rate of $\dot{M}=(3.0 \pm 2.1)\, \rm M_\odot \, yr^{-1}$ averaged over a time span of 40 Myr, which varies by a factor of a few over time-scales of ∼10 Myr. Most of the inflow appears to be consumed by star formation in the ring, which is currently occurring at a star formation rate (SFR) of $\simeq\!1.8\!-\!2 \, \rm M_\odot \, yr^{-1}$, suggesting that the inflow is causally controlling the SFR in the ring as a function of time.

 

ABSTRACT Analysis of the genes retained in the minimized Mycoplasma JCVI-Syn3A genome established that systems that repair or preempt metabolite damage are essential to life. Several genes known to have such functions were identified and experimentally validated, including 5-formyltetrahydrofolate cycloligase, coenzyme A (CoA) disulfide reductase, and certain hydrolases. Furthermore, we discovered that an enigmatic YqeK hydrolase domain fused to NadD has a novel proofreading function in NAD synthesis and could double as a MutT-like sanitizing enzyme for the nucleotide pool. Finally, we combined metabolomics and cheminformatics approaches to extend the core metabolic map of JCVI-Syn3A to include promiscuous enzymatic reactions and spontaneous side reactions. This extension revealed that several key metabolite damage control systems remain to be identified in JCVI-Syn3A, such as that for methylglyoxal. IMPORTANCE Metabolite damage and repair mechanisms are being increasingly recognized. We present here compelling genetic and biochemical evidence for the universal importance of these mechanisms by demonstrating that stripping a genome down to its barest essentials leaves metabolite damage control systems in place. Furthermore, our metabolomic and cheminformatic results point to the existence of a network of metabolite damage and damage control reactions that extends far beyond the corners of it that have been characterized so far. In sum, there can be little room left to doubt that metabolite damage and the systems that counter it are mainstream metabolic processes that cannot be separated from life itself. 

We use new HCN(1–0) data from the ACA Large-sample Mapping Of Nearby galaxies in Dense gas (ALMOND) survey to trace the kpc-scale molecular gas density structure and CO(2–1) data from the Physics at High Angular resolution in Nearby GalaxieS–Atacama Large Millimeter/submillimeter Array (PHANGS–ALMA) to trace the bulk molecular gas across 25 nearby star-forming galaxies. At 2.1 kpc scale, we measure the density-sensitive HCN/CO line ratio and the star formation rate (SFR)/HCN ratio to trace the star formation efficiency in the denser molecular medium. At 150 pc scale, we measure structural and dynamical properties of the molecular gas via CO(2–1) line emission, which is linked to the lower resolution data using an intensity-weighted averaging method. We find positive correlations (negative) of HCN/CO (SFR/HCN) with the surface density, the velocity dispersion, and the internal turbulent pressure of the molecular gas. These observed correlations agree with expected trends from turbulent models of star formation, which consider a single free-fall time gravitational collapse. Our results show that the kpc-scale HCN/CO line ratio is a powerful tool to trace the 150 pc scale average density distribution of the molecular clouds. Lastly, we find systematic variations of the SFR/HCN ratio with cloud-scale molecular gas properties, which are incompatible with a universal star formation efficiency. Overall, these findings show that mean molecular gas density, molecular cloud properties, and star formation are closely linked in a coherent way, and observations of density-sensitive molecular gas tracers are a useful tool to analyse these variations, linking molecular gas physics to stellar output across galaxy discs.

 

Abstract The CO-to-H 2 conversion factor ( α CO ) is critical to studying molecular gas and star formation in galaxies. The value of α CO has been found to vary within and between galaxies, but the specific environmental conditions that cause these variations are not fully understood. Previous observations on ~kiloparsec scales revealed low values of α CO in the centers of some barred spiral galaxies, including NGC 3351. We present new Atacama Large Millimeter/submillimeter Array Band 3, 6, and 7 observations of 12 CO, 13 CO, and C 18 O lines on 100 pc scales in the inner ∼2 kpc of NGC 3351. Using multiline radiative transfer modeling and a Bayesian likelihood analysis, we infer the H 2 density, kinetic temperature, CO column density per line width, and CO isotopologue abundances on a pixel-by-pixel basis. Our modeling implies the existence of a dominant gas component with a density of 2–3 × 10 3 cm −3 in the central ∼1 kpc and a high temperature of 30–60 K near the nucleus and near the contact points that connect to the bar-driven inflows. Assuming a CO/H 2 abundance of 3 × 10 −4 , our analysis yields α CO ∼ 0.5–2.0 M ⊙ (K km s −1 pc 2 ) −1 with a decreasing trend with galactocentric radius in the central ∼1 kpc. The inflows show a substantially lower α CO ≲ 0.1 M ⊙ (K km s −1 pc 2 ) −1 , likely due to lower optical depths caused by turbulence or shear in the inflows. Over the whole region, this gives an intensity-weighted α CO of ∼1.5 M ⊙ (K km s −1 pc 2 ) −1 , which is similar to previous dust-modeling-based results at kiloparsec scales. This suggests that low α CO on kiloparsec scales in the centers of some barred galaxies may be due to the contribution of low-optical-depth CO emission in bar-driven inflows. 

Abstract We compare mid-infrared (mid-IR), extinction-corrected H α , and CO (2–1) emission at 70–160 pc resolution in the first four PHANGS–JWST targets. We report correlation strengths, intensity ratios, and power-law fits relating emission in JWST’s F770W, F1000W, F1130W, and F2100W bands to CO and H α . At these scales, CO and H α each correlate strongly with mid-IR emission, and these correlations are each stronger than the one relating CO to H α emission. This reflects that mid-IR emission simultaneously acts as a dust column density tracer, leading to a good match with the molecular-gas-tracing CO, and as a heating tracer, leading to a good match with the H α . By combining mid-IR, CO, and H α at scales where the overall correlation between cold gas and star formation begins to break down, we are able to separate these two effects. We model the mid-IR above I ν = 0.5 MJy sr −1 at F770W, a cut designed to select regions where the molecular gas dominates the interstellar medium (ISM) mass. This bright emission can be described to first order by a model that combines a CO-tracing component and an H α -tracing component. The best-fitting models imply that ∼50% of the mid-IR flux arises from molecular gas heated by the diffuse interstellar radiation field, with the remaining ∼50% associated with bright, dusty star-forming regions. We discuss differences between the F770W, F1000W, and F1130W bands and the continuum-dominated F2100W band and suggest next steps for using the mid-IR as an ISM tracer. 

The processes of star formation and feedback, regulating the cycle of matter between gas and stars on the scales of giant molecular clouds (GMCs; ∼100 pc), play a major role in governing galaxy evolution. Measuring the time-scales of GMC evolution is important to identify and characterize the specific physical mechanisms that drive this transition. By applying a robust statistical method to high-resolution CO and narrow-band H α imaging from the PHANGS survey, we systematically measure the evolutionary timeline from molecular clouds to exposed young stellar regions on GMC scales, across the discs of an unprecedented sample of 54 star-forming main-sequence galaxies (excluding their unresolved centres). We find that clouds live for about 1−3 GMC turbulence crossing times (5−30 Myr) and are efficiently dispersed by stellar feedback within 1−5 Myr once the star-forming region becomes partially exposed, resulting in integrated star formation efficiencies of 1−8 per cent. These ranges reflect physical galaxy-to-galaxy variation. In order to evaluate whether galactic environment influences GMC evolution, we correlate our measurements with average properties of the GMCs and their local galactic environment. We find several strong correlations that can be physically understood, revealing a quantitative link between galactic-scale environmental properties and the small-scale GMC evolution. Notably, the measured CO-visible cloud lifetimes become shorter with decreasing galaxy mass, mostly due to the increasing presence of CO-dark molecular gas in such environment. Our results represent a first step towards a comprehensive picture of cloud assembly and dispersal, which requires further extension and refinement with tracers of the atomic gas, dust, and deeply embedded stars.