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The spectacular fossil fauna and flora preserved in the Upper Cretaceous terrestrial strata of North America’s Western Interior Basin record an exceptional peak in the diversification of fossil vertebrates in the Campanian, which has been termed the ‘zenith of dinosaur diversity’. The wide latitudinal distribution of rocks and fossils that represent this episode, spanning from northern Mexico to the northern slopes of Alaska, provides a unique opportunity to gain insights into dinosaur paleoecology and to address outstanding questions regarding faunal provinciality in connection to paleogeography and climate. Whereas reliable basin-wide correlations are fundamental to investigations of this sort, three decades of radioisotope geochronology of various vintages and limited compatibility has complicated correlation of distant fossil-bearing successions and given rise to contradictory paleobiogeographic and evolutionary hypotheses. Here we present new U–Pb geochronology by the CA-ID-TIMS method for 16 stratigraphically well constrained bentonite beds, ranging in age from 82.419 ± 0.074 Ma to 73.496 ± 0.039 Ma (2σ internal uncertainties), and the resulting Bayesian age models for six key fossil-bearing formations over a 1600 km latitudinal distance from northwest New Mexico, USA to southern Alberta, Canada. Our high-resolution chronostratigraphic framework for the upper Campanian of the Western Interior Basin reveals that despite their contrasting depositional settings and basin evolution histories, significant age overlap exists between the main fossil-bearing intervals of the Kaiparowits Formation (southern Utah), Judith River Formation (central Montana), Two Medicine Formation (western Montana) and Dinosaur Park Formation (southern Alberta). Pending more extensive paleontologic collecting that would allow more rigorous faunal analyses, our results support a first-order connection between paleoecologic and fossil diversities and help overcome the chronostratigraphic ambiguities that have impeded the testing of proposed models of latitudinal provinciality of dinosaur taxa during the Campanian.

 

Abstract The Late Campanian (Late Cretaceous), upper part of the El Disecado Member, El Gallo Formation, Baja California, México, preserves a rich fossil assemblage of microvertebrates and macrovertebrates, silicified logs, macroscopic plant remains, and pollen that was likely deposited as the distal part of a subaerial fan. The unit was episodic and high energy, with its salient features deriving from active river channels and sheet, debris-flow deposits. Landscape stability is indicated by the presence of compound paleosol horizons, containing Fe2O3 mottling in B horizons, cutans, and calcium carbonate concretions. All of these features indicate wet/dry cyclicity in subsurface horizons, likely attributable to such cyclicity in the climate. Drainage was largely to the north and to a lesser extent, the west; however, some current flow to the south and east is preserved which, in conjunction with the proximal location of marginal marine deposits, suggest the influence of tides in this setting. The fossil vertebrates preserved in this part of the El Disecado Member are almost exclusively allochthonous, preserved as disarticulated isolated clasts in hydraulic equivalence in the braided fluvial system. A relatively diverse microvertebrate assemblage is preserved, the largest components of which are first, dinosaurs, and second, turtles. Non-tetrapod fossils are relatively uncommon, perhaps reflecting an absence of permanent standing water in this depositional setting. Here we report a high-precision U-Pb date of 74.706 + 0.028 Ma (2σ internal uncertainty), obtained from zircons in an airfall tuff. The tuff is located low within the sequence studied; therefore, most of the sedimentology and fossils reported here are slightly younger. This date, which improves upon previously published 40Ar/39Ar geochronology, ultimately allows for comparison of these El Gallo faunas and environments with coeval ones globally. Primary stable isotopic nodules associated with roots in the paleosols of the terrestrial portion of the El Disecado Member are compared with ratios from similar sources from coeval northern and eastern localities in North America. Distinctive latitudinal gradients are observed in both δ13C and δ18O, reflecting the unique southern and western, coastal geographic position of this locality. These differences are best explained by differences in the floras that populated the northern and eastern localities, relative to the southern and western floras reported here.