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Flying snakes (genus Chrysopelea) glide without the use of wings. Instead, they splay their ribs and undulate through the air. A snake's ability to glide depends on how well its morphing wing-body produces lift and drag forces. However, previous kinematics experiments under-resolved the body, making it impossible to estimate the aerodynamic load on the animal or to quantify the different wing configurations throughout the glide. Here, we present new kinematic analyses of a previous glide experiment, and use the results to test a theoretical model of flying snake aerodynamics using previously measured lift and drag coefficients to estimate the aerodynamic forces. This analysis is enabled by new measurements of the center of mass motion based on experimental data. We found that quasi-steady aerodynamic theory under-predicts lift by 35% and over-predicts drag by 40%. We also quantified the relative spacing of the body as the snake translates through the air. In steep glides, the body is generally not positioned to experience tandem effects from wake interaction during the glide. These results suggest that unsteady 3D effects, with appreciable force enhancement, are important for snake flight. Future work can use the kinematics data presented herein to form test conditions for physical modeling, as well as computational studies to understand unsteady fluid dynamics effects on snake flight. 

Even the most simplified models of falling and gliding bodies exhibit rich nonlinear dynamical behavior. Taking a global view of the dynamics of one such model, we find an attracting invariant manifold that acts as the dominant organizing feature of trajectories in velocity space. This attracting manifold captures the final, slowly changing phase of every passive descent, providing a higher-dimensional analogue to the concept of terminal velocity, the terminal velocity manifold. Within the terminal velocity manifold in extended phase space, there is an equilibrium submanifold with equilibria of alternating stability type, with different stability basins. In this work, we present theoretical and numerical methods for approximating the terminal velocity manifold and discuss ways to approximate falling and gliding motion in terms of these underlying phase space structures. 

This paper introduces the trajectory divergence rate, a scalar field which locally gives the instantaneous attraction or repulsion of adjacent trajectories. This scalar field may be used to find highly attracting or repelling invariant manifolds, such as slow manifolds, to rapidly approximate hyperbolic Lagrangian coherent structures, or to provide the local stability of invariant manifolds. This work presents the derivation of the trajectory divergence rate and the related trajectory divergence ratio for two-dimensional systems, investigates their properties, shows their application to several example systems, and presents their extension to higher dimensions.