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Abstract An anelastic numerical model is used to study the influences of fine structure (FS) in the wind and stability profiles on gravity wave (GW) propagation in the Mesosphere and Lower Thermosphere (MLT). Large amplitude GWs interacting with FS, that is, thin regions of enhanced wind and stability, evolve very differently depending on the precise vorticity source and sink terms for small‐scale motions induced by the FS gradients. The resulting small‐scale dynamics are deterministic, promoting local instabilities, dissipation, and momentum deposition at locations and orientations determined by the initial FS. The resulting momentum depositions yield significant changes to the background wind structure, having scales and amplitudes comparable to the effects of large‐scale features in the ambient atmosphere. The deterministic nature of the large‐scale impacts further suggests that they can be estimated without fully resolving the underlying instability dynamics. Given the significant amplitudes and ubiquitous occurrence of FS throughout the atmosphere, the influences of these important and diverse flow evolutions merit inclusion in broader modeling efforts. 

New comparisons between the square of the generalized potential refractive index gradient M2, estimated from the very high-frequency (VHF) Middle and Upper Atmosphere (MU) Radar, located at Shigaraki, Japan, and unmanned aerial vehicle (UAV) measurements are presented. These comparisons were performed at unprecedented temporal and range resolutions (1–4 min and ⇠20 m, respectively) in the altitude range ⇠1.27–4.5 km from simultaneous and nearly collocated measurements made during the ShUREX (Shigaraki UAV-Radar Experiment) 2015 campaign. Seven consecutive UAV flights made during daytime on 7 June 2015 were used for this purpose. The MU Radar was operated in range imaging mode for improving the range resolution at vertical incidence (typically a few tens of meters). The proportionality of the radar echo power to M2 is reported for the first time at such high time and range resolutions for stratified conditions for which Fresnel scatter or a reflection mechanism is expected. In more complex features obtained for a range of turbulent layers generated by shear instabilities or associated with convective cloud cells, M2 estimated from UAV data does not reproduce observed radar echo power profiles. Proposed interpretations of this discrepancy are presented.