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Abstract Stream dissolved organic matter (DOM) is a globally important carbon flux and a locally important control on stream ecosystems, and therefore understanding controls on stream DOM fluxes and dynamics is crucial at both local and global scales. However, attributing process controls is challenging because both hydrological and biological controls on DOM are integrated and may vary over time and throughout stream networks. Our objective was to assess the patterns and corresponding controls of diel DOM cycles through a seasonal flow recession by using reach‐scale in situ sensors in a non‐perennial stream network. We identified five characteristic diel variations in DOM with differing phase and amplitude. During snowmelt flows, diel variations in DOM were consistent among sites and reflected diel flowpath shifts and photodegradation. Evapotranspiration‐driven diel stage oscillations emerged at two upstream sites, shaping diel DOM patterns indirectly, by creating conditions for instream DOM processing. At a spring‐fed site, minimal diel variation was observed throughout the summer whereas at an intermittent reach, daily drying and rewetting created biogeochemical hot moments. This research demonstrates that controls on DOM vary over time and space, even in close proximity, generating asynchronous fDOM patterns during low flows, illuminating shifts in biogeochemical processes and flowpaths. 

Abstract In the western United States, water supplies largely originate as snowmelt from forested land. Forests impact the water balance of these headwater streams, yet most predictive runoff models do not explicitly account for changing snow‐vegetation dynamics. Here, we present a case study showing how warmer temperatures and changing forests in the Henrys Fork of the Snake River, a seasonally snow‐covered headwater basin in the Greater Yellowstone Ecosystem, have altered the relationship between April 1st snow water equivalent (SWE) and summer streamflow. Since the onset and recovery of severe drought in the early 2000s, predictive models based on pre‐drought relationships over‐predict summer runoff in all three headwater tributaries of the Henrys Fork, despite minimal changes in precipitation or snow accumulation. Compared with the pre‐drought period, late springs and summers (May–September) are warmer and vegetation is greener with denser forests due to recovery from multiple historical disturbances. Shifts in the alignment of snowmelt and energy availability due to warmer temperatures may reduce runoff efficiency by changing the amount of precipitation that goes to evapotranspiration versus runoff and recharge. To quantify the alignment between snowmelt and energy on a timeframe needed for predictive models, we propose a new metric, the Vegetation‐Water Alignment Index (VWA), to characterize the synchrony of vegetation greenness and snowmelt and rain inputs. New predictive models show that in addition to April 1st SWE, the previous year's VWA and summer reference evapotranspiration are the most significant predictors of runoff in each watershed and provide more predictive power than traditionally used metrics. These results suggest that the timing of snowmelt relative to the start of the growing season affects not only annual partitioning of streamflow, but can also determine the groundwater storage state that dictates runoff efficiency the following spring. 

Abstract Non‐perennial streams are receiving increased attention from researchers, however, suitable methods for measuring their hydrologic connectivity remain scarce. To address this deficiency, we developed Bayesian statistical approaches for measuring both average active stream length, and a new metric called average communication distance. Average communication distance is a theoretical increasedeffective distancethat stream‐borne materials must travel, given non‐continuous streamflow. Because it is the product of the inverse probability of surface water presence and stream length, the average communication distance of a non‐perennial stream segment will be greater than its actual physical length. As an application we considered Murphy Creek, a simple non‐perennial stream network in southwestern Idaho, USA. We used surface water presence/absence data obtained in 2019, and priors for the probability of surface water, based on predictions from an existing regional United States Geological Survey model. Average communication distance posterior distributions revealed locations where effective stream lengths increased dramatically due to flow rarity. We also found strong seasonal (spring, summer, fall) differences in network‐level posterior distributions of both average stream length and average communication distance. Our work demonstrates the unique perspectives concerning network drying provided by communication distance, and demonstrates the general usefulness of Bayesian approaches in the analysis of non‐perennial streams. 

Abstract As drought and wildfire frequency increase across the western United States, our ability to predict how water resources will respond to these disturbances depends on our understanding of the feedbacks that maintain watershed function and streamflow. Previous studies of non‐perennial headwater streams have ranked drivers of low‐flow conditions; however, there is a limited understanding of the interactions between these drivers and the processes through which these interactions affect streamflow. Here, we use stream water level, soil moisture, sap flow, and vapor pressure deficit data to investigate ecohydrological interactions along a mountainous headwater stream. Correlation and cross‐correlation analyses of these variables show that ecohydrological interactions are (a) nonlinear and (b) interconnected, suggesting that analyses assuming linearity and independence of each driver are inadequate for quantifying these interactions. To account for these issues and investigate causal linkages, we use convergent cross‐mapping (CCM) to characterize the feedbacks that influence non‐perennial streamflow. CCM is a nonlinear, dynamic method that has only recently been applied to hydrologic systems. CCM results reveal that atmospheric losses associated with local sap flow and vapor pressure deficit are driving changes in soil moisture and streamflow (p < 0.01) and that atmospheric losses influence stream water more directly than shallow soil moisture. These results also demonstrate that riparian processes continue to affect subsurface flows in the channel corridor even after stream drying. This study proposes a nonlinear framework for quantifying the ecohydrologic interactions that may determine how headwater streams respond to disturbance. 

Abstract Intermittent streams currently constitute >50% of the global river network, and the number of intermittent streams is expected to increase due to changes in land use and climate. Surface flows are known to expand and contract within the headwater channel network due to changes in the water table driven by climate, often changing seasonally. However, the underlying causes of disconnections and reconnections throughout the stream network remain poorly understood and may reflect subsurface flow capacity. We assess how 3D subsurface flowpaths control local surface flows at Gibson Jack Creek in the Rocky Mountains, Idaho, USA. Water table dynamics, hydraulic gradients, and hyporheic exchange were monitored along a 200‐m section of the stream throughout the seasonal recession in WY2018. Shallow lateral hillslope‐riparian‐stream connectivity was more frequent in transects spanning perennially flowing stream reaches than intermittent reaches. During low‐flow periods, larger losing vertical hydraulic gradients were observed in paired piezometers in intermittent reaches than in adjacent perennial reaches. Contrary to dominant conceptual models, longitudinal measurements of hydrologic exchange in both intermittent and perennial reaches were seasonally variable except for one perennial reach that showed consistent significant gains. Observed drying dynamics, as well as subsurface pathways, were highly variable even over short distances (30 m). Flow probability and subsurface flow capacity at upstream locations can be assessed with an outlet hydrograph and upstream flow measurements. Accurate characterization of subsurface storage, discharge, and connection is critical to understanding the drivers of drying cycles in intermittent streams and their likely responses to future change. 

Abstract Over half of global rivers and streams lack perennial flow, and understanding the distribution and drivers of their flow regimes is critical for understanding their hydrologic, biogeochemical, and ecological functions. We analyzed nonperennial flow regimes using 540 U.S. Geological Survey watersheds across the contiguous United States from 1979 to 2018. Multivariate analyses revealed regional differences in no‐flow fraction, date of first no flow, and duration of the dry‐down period, with further divergence between natural and human‐altered watersheds. Aridity was a primary driver of no‐flow metrics at the continental scale, while unique combinations of climatic, physiographic and anthropogenic drivers emerged at regional scales. Dry‐down duration showed stronger associations with nonclimate drivers compared to no‐flow fraction and timing. Although the sparse distribution of nonperennial gages limits our understanding of such streams, the watersheds examined here suggest the important role of aridity and land cover change in modulating future stream drying. 

Abstract Dissolved organic carbon (DOC) concentrations vary among headwaters, with variation typically decreasing with watershed area. We hypothesized that streamflow intermittence could be an important source of variation in DOC concentrations across a small watershed, through (a) temporal legacies of drying on organic matter accumulation and biotic communities and (b) spatial patterns of connectivity with DOC sources. To test these hypotheses, we conducted three synoptic water chemistry sampling campaigns across a 25.5‐km²watershed in south‐eastern Idaho during early spring, late summer, and late fall. Using changepoint analysis, we found that DOC variability collapsed at a consistent location (watershed areas ~1.3 to ~1.8 km²) across seasons, which coincided with the watershed area where variability in streamflow intermittence collapsed (~1.5 km²). To test hypothesized mechanisms through which intermittence may affect DOC, we developed temporal, spatial, and spatio‐temporal metrics of streamflow intermittence and related these to DOC concentrations. Streamflow intermittence was a strong predictor of DOC across seasons, but different metrics predicted DOC depending on season. Seasonal changes in the effects of intermittence on DOC reflected seasonal changes from instream to flowpath controls. A metric that captured spatial connectivity to sources significantly predicted DOC during high flows, when DOC is typically controlled by transport. In contrast, a reach‐scale temporal metric of intermittence predicted DOC during the late growing season, when DOC is typically controlled by instream processes and when legacy effects of drying (e.g., diminished biological communities) would likely affect DOC. The effects of intermittence on DOC extend beyond temporal legacies at a point. Our results suggest that legacy effects of intermittence do not propagate downstream in this system. Instead, snapshots of spatial patterns of intermittence upstream of a reach are critical for understanding spatial patterns of DOC through connectivity to DOC sources, and these processes drive patterns of DOC even in perennial reaches.