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Abstract Collectively locomoting animals are often viewed as analogous to states of matter in that group-level phenomena emerge from individual-level interactions. Applying this framework to fish schools and bird flocks must account for visco-inertial flows as mediators of the physical interactions. Motivated by linear flight formations, here we show that pairwise flow interactions tend to promote crystalline or lattice-like arrangements, but such order is disrupted by unstably growing positional waves. Using robotic experiments on “mock flocks” of flapping wings in forward flight, we find that followers tend to lock into position behind a leader, but larger groups display flow-induced oscillatory modes – “flonons” – that grow in amplitude down the group and cause collisions. Force measurements and applied perturbations inform a wake interaction model that explains the self-ordering as mediated by spring-like forces and the self-amplification of disturbances as a resonance cascade. We further show that larger groups may be stabilized by introducing variability among individuals, which induces positional disorder while suppressing flonon amplification. These results derive from generic features including locomotor-flow phasing and nonreciprocal interactions with memory, and hence these phenomena may arise more generally in macroscale, flow-mediated collectives. 

Significance Fish and birds moving in groups are thought to benefit from hydrodynamic or aerodynamic interactions between individuals. To better understand these effects, we devise a robotic “school” of flapping swimmers whose formations and motions come about from flow interactions. Surprisingly, we find that the flows naturally generated during swimming can also prevent collisions and separations, allowing even uncoordinated individuals with different flapping motions to travel together. Other benefits include freeloading by a “lazy” follower who keeps up with a faster-flapping leader by surfing on its wake. More generally, our study provides complete maps linking flapping motions to group locomotion, which is needed to test whether flow interactions are also exploited by animals. 

The passive flight of a thin wing or plate is an archetypal problem in flow–structure interactions at intermediate Reynolds numbers. This seemingly simple aerodynamic system displays an impressive variety of steady and unsteady motions that are familiar from fluttering leaves, tumbling seeds and gliding paper planes. Here, we explore the space of flight behaviours using a nonlinear dynamical model rooted in a quasisteady description of the fluid forces. Efficient characterisation is achieved by identification of the key dimensionless parameters, assessment of the steady equilibrium states and linear analysis of their stability. The structure and organisation of the stable and unstable flight equilibria proves to be complex, and seemingly related factors such as mass and buoyancy-corrected weight play distinct roles in determining the eventual flight patterns. The nonlinear model successfully reproduces previously documented unsteady states such as fluttering and tumbling while also predicting new types of motions, and the linear analysis accurately accounts for the stability of steady states such as gliding and diving. While the conditions for dynamic stability seem to lack tidy formulae that apply universally, we identify relations that hold in certain regimes and which offer mechanistic interpretations. The generality of the model and the richness of its solution space suggest implications for small-scale aerodynamics and related applications in biological and robotic flight. 

Mechanical systems with moving points of contact—including rolling, sliding, and impacts—are common in engineering applications and everyday experiences. The challenges in analyzing such systems are compounded when an object dynamically explores the complex surface shape of a moving structure, as arises in familiar but poorly understood contexts such as hula hooping. We study this activity as a unique form of mechanical levitation against gravity and identify the conditions required for the stable suspension of an object rolling around a gyrating body. We combine robotic experiments involving hoops twirling on surfaces of various geometries and a model that links the motions and shape to the contact forces generated. The in-plane motions of the hoop involve synchronization to the body gyration that is shown to require damping and sufficiently high launching speed. Further, vertical equilibrium is achieved only for bodies with “hips” or a critical slope of the surface, while stability requires an hourglass shape with a “waist” and whose curvature exceeds a critical value. Analysis of the model reveals dimensionless factors that successfully organize and unify observations across a wide range of geometries and kinematics. By revealing and explaining the mechanics of hula hoop levitation, these results motivate strategies for motion control via geometry-dependent contact forces and for accurately predicting the resulting equilibria and their stability. 

Extensive studies of the hydraulics of pipes have focused on limiting cases, such as fully-developed laminar or turbulent flow through long conduits and the accelerating flow through an orifice, for which there exist laws relating pressure drop and flow rate. We carry out experiments on smooth, circular pipes for dimensions and flow rates that interrogate intermediate conditions between the well-studied limits. Organizing this information in terms of dimensionless friction factor, Reynolds number and pipe aspect ratio yields a surface $$f_D(Re,\alpha )$$ that is shown to match the three laws associated with developed laminar, developed turbulent, and orifice flows. While each law fails outside its applicable range of $$(Re,\alpha )$$ , we present a hybrid theoretical–empirical model that includes inlet, development and transition effects, and that proves accurate to approximately 10 % over wide ranges of $Re$ and $$\alpha$$ . We also present simple formulas for the boundaries between the three hydraulic regimes, which intersect at a triple point. Measurements show that sipping through a straw is an everyday example of such intermediate conditions not accounted for by existing laws but described accurately by our model. More generally, our findings provide formulas for predicting frictional resistance for intermediate- $Re$ flows through finite-length pipes.