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Gas flows between galaxies and the circumgalactic medium (CGM) play a central role in galaxy evolution and can become observable as giant nebulae when illuminated by the quasars. We present an ensemble study of nebulae around 30 UV-luminous quasars at z=0.4-1.4 from the CUBS and MUSEQuBES surveys, 27 of which are detected in extended [O II] and/or [O III] emission. Based on a joint analysis of nebular morphology and surrounding galaxy environments, we introduce three morpho-kinematic classifications. We identify eleven irregular, large-scale (>50 kpc) systems, many of which are likely interaction-related; twelve compact host-galaxy-scale nebula, likely tracing CGM/ISM gas; and four systems with complex morphologies of uncertain origin. We introduce a quantitative measure of the spatial and kinematic association between nebulae and quasar-host group galaxies, finding a statistically significant association for ten nebulae, most of which are irregular, large-scale nebulae, consistent with qualitative analysis. Radio jets are detected in six systems, with no strong correlation found between radio activity and nebular emission. The [O II] nebulae are more asymmetric than their Ly α counterparts at z>2 , but bear more similarity to H I gas observed in 21 cm around local elliptical galaxies. Blueshifted-redshifted patterns, likely tracing gas rotation, are observed in roughly 30% of the systems, though disturbed kinematics suggest that feedback may also be important. These results show that giant quasar nebulae are not a uniform class of objects, but instead arise through multiple pathways shaped by host-galaxy gas, galaxy interactions, group environments, and quasar activity, with the most striking cases associated with galaxy interactions. 

Abstract Flavin-dependent halogenases (FDHs) are a class of enzymes renowned for their regioselective ability to precisely insert halogen atoms into small aromatic compounds. Halogen incorporation can enhance the physicochemical and biological properties of molecules, making them valuable for agrochemical and pharmaceutical applications. Through bioinformatic mining of bacterial genomes, we discovered and functionally characterized SnFDHal, an efficient tryptophan 5-halogenase fromStreptomyces nourseiNRRL B-1714. This halogenase operates across a broad pH range and exhibits a melting temperature of 46.7 °C at both pH 6 and pH 8, which is comparable to thermophilic halogenases such as Th-Hal and BorH, and notably higher than those of mesophilic counterparts. Steady-state kinetic analysis revealed that SnFDHal displays superior catalytic efficiency for the chlorination of L-tryptophan compared to other FDHs reported to date. Structural modeling of its active site suggests that a conserved bulky phenylalanine residue (F49) promotes halogenation at the C-5 position of L-tryptophan, consistent with experimental findings. The combination of high catalytic efficiency and thermostability positions SnFDHal as a promising biocatalyst for applications in agrochemical and pharmaceutical industries. 

Protein tyrosine phosphatases (PTPs) are a family of enzymes that play important roles in regulating cellular signaling pathways. The activity of these enzymes is regulated by the motion of a catalytic loop that places a critical conserved aspartic acid side chain into the active site for acid–base catalysis upon loop closure. These enzymes also have a conserved phosphate-binding loop that is typically highly rigid and forms a well-defined anion-binding nest. The intimate links between loop dynamics and chemistry in these enzymes make PTPs an excellent model system for understanding the role of loop dynamics in protein function and evolution. In this context, archaeal PTPs, which have often evolved in extremophilic organisms, are highly understudied, despite their unusual biophysical properties. We present here an engineered chimeric PTP (ShufPTP) generated by shuffling the amino acid sequence of five extant hyperthermophilic archaeal PTPs. Despite ShufPTP’s high sequence similarity to its natural counterparts, it presents a suite of unique properties, including high flexibility of the phosphate binding P-loop, facile oxidation of the active-site cysteine, mechanistic promiscuity, and, most notably, hyperthermostability, with a denaturation temperature likely >130 °C (>8 °C higher than the highest recorded growth temperature of any archaeal strain). Our combined structural, biochemical, biophysical, and computational analysis provides insight both into how small steps in evolutionary space can radically modulate the biophysical properties of an enzyme and showcases the tremendous potential of archaeal enzymes for biotechnology, to generate novel enzymes capable of operating under extreme conditions. 

Abstract Feedback from active galactic nuclei (AGN) is widely acknowledged to regulate the growth of massive galaxies, though its driving mechanisms are debated. Prevailing theories suggest that AGN-driven outflows are driven either by radiation pressure acting directly on the dusty interstellar medium (ISM) or by hot winds entraining cooler ISM gas, but the relative contribution of each mechanism remains uncertain. By combining optical emission line measurements with highly ionized UV emission lines, it is possible to constrain whether the pressure source applied to ionized clouds is primarily radiation or primarily hydrodynamic, and thus constrain the dominant driver. This study presents the first multiobject analysis of far-UV spectra from galactic-scale AGN-driven outflows in obscured quasars, based on Cosmic Origins Spectrograph observations of five low-redshift targets. By comparing narrow-line region UV emission-line ratios to theoretical models that vary the importance of the two pressure sources, we find that three out of five targets fall within the radiation-pressure-dominated regime. A fourth target exhibits intermediate emission-line ratios that suggest radiation pressure and pressure from a hot wind are both dynamically important. Finally, the lowest-luminosity object in our sample may have a dynamically important hot wind component, but nondetections prevent a clear conclusion in this case. These results suggest radiation pressure dominates circumnuclear narrow-line region cloud dynamics, but pressure from a hot wind also plays a role in some cases. This is consistent with AGN feedback scenarios mediated by radiation pressure or a short-lived hot wind phase that dissipates after initially accelerating outflows. 

Abstract In dwarf galaxy models, outflows expel metal-enriched interstellar medium (ISM) into the circumgalactic medium (CGM) to reproduce their observed low metallicities, but measurements of dwarf CGM properties are scarce. We present a study of the CGM of an isolated dwarf atz= 0.5723 with a stellar mass of ≈5 × 10⁷M_⊙and star formation rate (≈0.05M_⊙yr⁻¹) and ISM metallicity ([O/H] ≈ −0.9) consistent with the star-forming main sequence and mass–metallicity relation. A background quasar sight line with archival UV spectra probes the dwarf’s CGM at a projected distance of 28 kpc, corresponding to approximately half of the estimated virial radius. The dwarf’s CGM is detected in Hi;intermediate metal ions of Ciii, Oiii, Oiv, and Sv; and kinematically broader, highly ionized Ovibut is undetected in Nivand Neviii. Photoionization modeling of the intermediate ions indicates a modest volume filling factor (∼6% along the sight line or ∼2% globally) and a mass of ∼2 × 10⁸M_⊙, ∼4× higher than the dwarf’s stellar mass but ∼10× less than the highly ionized CGM. The Ovikinematics are comparable to the dwarf’s estimated virial velocity, suggesting that it is likely associated with cool, photoionized, and volume-filling CGM, with bulk motion or turbulence dominating over thermal pressure. The metallicity inferred for the intermediate ions is [O/H] = −0.6, but with low relative abundances of [C/O] = −0.6 and [N/O] < −1.0. The [N/O] is below levels expected of the dwarf’s ISM but consistent with core-collapse supernova ejecta, suggesting that supernova-enriched gas escaped the dwarf without mixing significantly with ISM enriched in nitrogen from evolved, low-mass stars. 

Abstract We report statistically significant detection of Hi21 cm emission from intermediate-redshift (z ≈ 0.2–0.6) galaxies. By leveraging multisightline galaxy survey data from the Cosmic Ultraviolet Baryon Survey and deep radio observations from the MeerKAT Absorption Line Survey, we have established a sample of ≈6000 spectroscopically identified galaxies in 11 distinct fields to constrain the neutral gas content at intermediate redshifts. The galaxies sample a broad range in stellar mass, from $\log M_{\mathrm{star}}/M_{\odot }\approx 8$ to $\log M_{\mathrm{star}}/M_{\odot }\approx 11$ , with a median of ${〈\log M_{\mathrm{star}}/M_{\odot }〉}_{\mathrm{med}}\approx 10$ and a wide range in redshift fromz ≈ 0.24 toz ≈ 0.63 with a median of 〈z〉_med = 0.44. While no individual galaxies show detectable Hiemission, the emission line signal is detected in the stacked spectra of all subsamples at greater than 4σsignificance. The observed total Hi21 cm line flux translates to a Himass,M_{H I}≈10¹⁰M_⊙. We find a high Hi-to-stellar-mass ratio ofM_HI/M_star ≈ 6 for low-mass galaxies with $〈\log M_{\mathrm{star}}/M_{\odot }〉\approx 9.3$ (>3.7σ). For galaxies with $〈\log M_{\mathrm{star}}/M_{\odot }〉\approx 10.6$ , we findM_HI/M_star ≈ 0.3 (>4.7σ). In addition, the redshift evolution of Himass, 〈M_{H I}〉, in both low- and high-mass field galaxies, inferred from the stacked emission-line signal, aligns well with the expectation from the cosmic star formation history. This suggests that the overall decline in the cosmic star formation activity across the general galaxy population may be connected to a decreasing supply of neutral hydrogen. Finally, our analysis has revealed significant 21 cm signals at distances greater than 75 kpc from these intermediate-redshift galaxies, indicating a substantial reservoir of Higas in their extended surroundings. 

Abstract The Ovi1032, 1038 Å line is a key probe of cooling gas in the circumgalactic medium (CGM) of galaxies but has been observed to date primarily in absorption along single sight lines. We present deep Hubble Space Telescope (HST) Solar Blind Channel of the Advanced Camera for Surveys observations of the compact, massive starburst Makani. Makani hosts a 100 kpc, [Oii]-emitting galactic wind driven by two episodes of star formation over 400 Myr. We detect Oviand Lyαemission across the [Oii] nebula with similar morphology and extent, out tor≈ 50 kpc. Using differential narrowband imaging, we separate Lyαand Oviand show that the Oviemission is comparable in brightness to [Oii], withL_{O VI}= 4 × 10⁴²erg s⁻¹. The similar hourglass morphology and size of [Oii] and Oviimplicate radiative cooling atT= 10^5.5K in a hot–cold interface. This may occur as theT> 10⁷K CGM—or the hot fluid driving the wind—exchanges mass with theT≈ 10⁴K clouds entrained in (or formed by) the wind. The optical/UV line ratios may be consistent with shock ionization, although uncertain attenuation and Lyαradiative transfer complicate the interpretation. The detection of Oviin Makani lies at the bleeding edge of the UV imaging capabilities of HST and provides a benchmark for future emission-line imaging of the CGM with a wide-area UV telescope. 

Abstract The shallow potential wells of star-forming dwarf galaxies make their surrounding circumgalactic and intergalactic medium (CGM/IGM) sensitive laboratories for studying the inflows and outflows thought to regulate galaxy evolution. We present new absorption-line measurements in quasar sight lines, probing within projected distances of <300 kpc from 91 star-forming field dwarf galaxies with a median stellar mass of $\log M_{\star }/M_{\odot }\approx 8.3$ at 0.077 <z< 0.73, from the Cosmic Ultraviolet Baryon Survey (CUBS). In this redshift range, the CUBS quasar spectra cover a suite of transitions including Hi, low, and intermediate metal ions (e.g., Cii, Siii, Ciii, and Siiii), and highly ionized Ovi. This CUBS-Dwarfs survey enables constraints with samples nine times larger than past dwarf CGM/IGM studies with similar ionic coverage. We find that low and intermediate ionization metal absorption is rare around dwarf galaxies, consistent with previous surveys of local dwarfs. In contrast, highly ionized Oviis commonly observed in sight lines that pass within the virial radius of a dwarf, and Ovidetection rates are nonnegligible at projected distances of 1−2× the virial radius. Based on these measurements, we estimate that the Ovi-bearing phase of the CGM/IGM accounts for a dominant share of the metal budget of dwarf galaxies. The absorption kinematics suggest that a relatively modest fraction of the Ovi-bearing gas is formally unbound. Together, these results imply that low-mass systems atz≲ 1 effectively retain a substantial fraction of their metals within the nearby CGM and IGM. 

Abstract This paper presents a newly established sample of 103 unique galaxies or galaxy groups at 0.4 ≲z≲ 0.7 from the Cosmic Ultraviolet Baryon Survey (CUBS) for studying the warm-hot circumgalactic medium (CGM) probed by both Oviand Neviiiabsorption. The galaxies and associated neighbors are identified at <1 physical Mpc from the sightlines toward 15 CUBS QSOs atz_QSO≳ 0.8. A total of 30 galaxies or galaxy groups exhibit associated Oviλλ1031, 1037 doublet absorption within a line-of-sight velocity interval of ±250 km s⁻¹, while the rest show no trace of Ovito a detection limit of $\log N_{\mathrm{OVI}}/{\mathrm{cm}}^{-2}\approx 13.7$ . Meanwhile, only five galaxies or galaxy groups exhibit the Neviiiλλ770, 780 doublet absorption, down to a limiting column density of $\log N_{\mathrm{NeVIII}}/{\mathrm{cm}}^{-2}\approx 14.0$ . These Ovi- and Neviii-bearing halos reside in different galaxy environments with stellar masses ranging from $\log M_{\mathrm{star}}/M_{\odot }\approx 8$ to ≈11.5. The warm-hot CGM around galaxies of different stellar masses and star formation rates exhibits different spatial profiles and kinematics. In particular, star-forming galaxies with $\log M_{\mathrm{star}}/M_{\odot }\approx 9–11$ show a significant concentration of metal-enriched warm-hot CGM within the virial radius, while massive quiescent galaxies exhibit flatter radial profiles of both column densities and covering fractions. In addition, the velocity dispersion of Oviabsorption is broad withσ_υ> 40 km s⁻¹for galaxies of $\log M_{\mathrm{star}}/M_{\odot }>9$ within the virial radius, suggesting a more dynamic warm-hot halo around these galaxies. Finally, the warm-hot CGM probed by Oviand Neviiiis suggested to be the dominant phase in sub-L* galaxies with $\log M_{\mathrm{star}}/M_{\odot }\approx 9–10$ based on their high ionization fractions in the CGM. 

Abstract The formation of the first supermassive black holes is expected to have occurred in some most pronounced matter and galaxy overdensities in the early universe. We have conducted a submillimeter wavelength continuum survey of 54z∼ 6 quasars using the Submillimeter Common-User Bolometre Array-2 on the James Clerk Maxwell Telescope to study the environments aroundz∼ 6 quasars. We identified 170 submillimeter galaxies (SMGs) with above 3.5σdetections in 450 or 850μm maps. Their far-IR luminosities are (2.2–6.4) × 10¹²L_⊙, and their star formation rates are ∼400–1200M_⊙yr⁻¹. We also calculated the SMGs’ differential and cumulative number counts in a combined area of ∼620 arcmin². To a 4σdetection (at ∼5.5 mJy), SMGs’ overdensity is ${0.68}_{-0.19}^{+0.21}$ (±0.19), exceeding the blank-field source counts by a factor of 1.68. We find that 13/54 quasars show overdensities (at ∼5.5 mJy) ofδ_SMG∼ 1.5–5.4. The combined area of these 13 quasars exceeds the blank-field counts with the overdensity to 5.5 mJy ofδ_SMG∼ ${2.46}_{-0.55}^{+0.64}$ (±0.25) in the regions of ∼150 arcmin². However, the excess is insignificant on the bright end (e.g., 7.5 mJy). We also compare results with previous environmental studies of Lyαemitters and Lyman break galaxies on a similar scale. Our survey presents the first systematic study of the environment of quasars atz∼ 6. The newly discovered SMGs provide essential candidates for follow-up spectroscopic observations to test whether they reside in the same large-scale structures as the quasars and search for protoclusters at an early epoch.