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The Norian–Rhaetian boundary (Late Triassic) represents an important precursor extinction event to the end- Triassic mass extinction, but the biotic and geochemical shifts are not well-understood due to poor stratigraphic constraints. Here we examine the microfossil record for metazoans and protists on a Panthalassan carbonate ramp (Gabbs Formation, Nevada, U.S.A.) during the late Norian to mid-Rhaetian, and correlate changes in these assemblages with macrofossil shifts and geochemical data (strontium and carbon isotopes). In the latest Norian, demosponge spicules represent a small proportion of shallow marine biosediments. Demosponges are joined in the earliest Rhaetian by increasingly abundant hypersilicified sponge spicules and silica-limited hexactinellid sponge spicules synchronous with a negative strontium isotope excursion indicating increased hydrothermal or volcanic activity. Common carbonate microfossils such as echinoderm ossicles and ostracods are typically silicified in these deposits as well, suggesting increased silicic porewater. The source for increased dissolved silica in shallow marine systems is suggested to be hydrothermal vent degassing, likely associated with increased tectonic rifting activity. Mid-Rhaetian microfossil assemblages exhibit evidence for intermitted anoxia in reducing conditions, supporting a scenario in which environmental stress was a prolonged feature of much of the Rhaetian Stage, rather than a short-term event in the terminal Rhaetian. While there is no marine sedimentary record of volcanism recognized for this interval, biosedimentary assemblages may serve as proxies for geochemical conditions associated with rifting. 

The Late Triassic was a time of major evolutionary transition for marine vertebrates, with the emergence of important new clades and the expansion of durophagy. The diversity record of shallow marine vertebrates has been limited by poor preservation, and palaeogeographic ranges of common marine vertebrates is poorly constrained. Isolated ichthyoliths are an important resource for documenting taxonomic and ecological diversity in shallow marine environments and have a relatively high preservation potential. Here, we report the first Upper Triassic elasmobranchs, neopterygians and chondrosteans from Nevada, and contribute 14 new generic occurrences. The vertebrates represented in this survey include teeth of specialised durophages and piscivores, and a tentative reconstruction of the shallow marine ecosystem trophic niches based on dietary interpretations and known shelly macrofossils from this locality. The ichthyolith assemblage exhibited greater similarity to low-latitude Europe than high-latitude Canada localities, and the results of this study suggest that several taxa previously considered to be endemic to Europe may be cosmopolitan. The present study highlights the need for additional research in early Mesozoic ichthyoliths in order to establish diversity patterns, paleobiogeographic ranges, and timing of adaptive radiations among key groups of marine vertebrates in the Triassic Period in suboptimal preservation conditions. 

Sections of the Gabbs Formation exposed near New York Canyon, Nevada, have long been recognized as important sites for Late Triassic and Early Jurassic stratigraphy, and the Norian-Rhaetian parts of these sections continue to be important for defining this boundary (NRB). The two candidate sections for the base of the Rhaetian are in Tethys; both sections utilize the first occurrence of the conodont species Misikella posthernsteini as a proxy for the boundary. Although not a candidate section, data from New York Canyon will help to determine the most suitable position for the NRB, especially in Panthalassa. Previous reports of conodonts from New York Canyon recognized a fauna with Mockina englandi, Mo. bidentata and morphotypes of Mo. mosheri in the Nun Mine Member, below isolated occurrences of Zieglericonus rhaeticum and Mi. posthernsteini in the Mount Hyatt and Muller Canyon members. The first occurrence of Mi. posthernsteini in the section occurs well above the first occurrence of Rhaetian ammonoids (Paracochloceras amoenum) and together with late Rhaetian radiolarians. It is also above excursions in Sr- and C-isotopes, both of which correlate with Tethyan NRB excursions. Therefore, the NRB has previously been placed much lower in North America than Tethys, at the first occurrences of the radiolarian Proparvicingula moniliformis and the conodont Mo. mosheri morphotype C. To help reconcile the biochronological and geochemical data from New York Canyon, new conodont samples have been collected from the Nun Mine and Mt Hyatt members at the New York Canyon Road and Luning Draw sections. These samples contain: Mo. englandi, Mo. bidentata, and Mo. mosheri morphotypes B and C, all previously reported from New York Canyon, although this is the first record of Mo. mosheri morphotype C from the Nun Mine Member; Parvigondolella spp. B and C, from much lower in the Nun Mine Member than previously reported; and Pa. andrusovi, which has not previously been recorded from North America. Overall, this fauna represents the Mo. bidentata and Mo. mosheri zones of North America, equivalent to the Sevatian Mo. bidentata and Pa. andrusovi zones of Tethys. This would be consistent with a higher placement of the NRB at New York Canyon; however, if the NRB is to be recognized at the first occurrence of Mo. mosheri morphotype C, then the boundary must be lower than previously thought, within the Nun Mine Member.