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High latitude upper atmospheric inter‐hemispheric asymmetry (IHA) tends to be enhanced during geomagnetic storms, which may be due to the complex spatiotemporal changes and magnitude modifications in field aligned currents (FACs) and particle precipitation (PP). However, the relative contribution of FACs and PP to IHA in high‐latitude forcing and energy is not well understood. The IHA during the 2015 St. Patrick’s Day storm has been investigated using the global ionosphere thermosphere model (GITM), driven by FACs from the Active Magnetosphere and Planetary Electrodynamics Response Experiment (AMPERE) and PP from the Assimilative Mapping of Ionospheric Electrodynamics (AMIE). A comprehensive study of the (a) relative contributions of FACs and PP to electric potential and Joule heating and (b) sensitivity of electric potential and Joule heating to the changes in magnitude and distribution of FACs and PP is presented. The results indicate that FACs lead to larger potential and Joule heating changes compared with PP. The spatial variations of potential and Joule heating are also affected by variation in FACs. As for asymmetric magnitude and distribution, it is found that electric potential and Joule heating are more sensitive to changes in the distribution of FACs and PP than the magnitude of FACs and PP. A new spatial asymmetry index (SAI) is introduced, which reveals spatial asymmetric details that are often overlooked by previous studies. This sensitivity study reveals the relative contributions in high‐latitude forcing and emphasizes the importance of obtaining accurate FACs and PP in both hemispheres.

We present a number of unique observations of ionospheric anomalies following the Hunga‐Tonga Hunga‐Ha'apai (HTHH) volcanic eruption on 15 January 2022. All are based on non‐dedicated geodetic satellite systems: Global Positioning System tracking of Low Earth Orbit (LEO) CubeSats, intersatellite tracking between two GRACE Follow‐On satellites, satellite radar altimeters to the ocean surface, and Doppler radio beacons from ground stations to LEO geodetic satellites. Their observations revealed the development of anomalously large trough‐like plasma depletions, along with plasma bubbles, in the equatorial regions of the Pacific and East Asian sectors. Trough‐like plasma depletions appeared to be confined within approximately ±20° magnetic latitude, accompanied by density enhancements just outside this latitude range. These plasma depletions and enhancements were aligned with the magnetic equator and occurred across broad longitudes. They were detected in regions where atmospheric waves from the HTHH eruption passed through around the time of the sunset terminator. We interpret these phenomena in terms of theEdynamo electric fields driven by atmospheric waves from the eruption. The uplift of the ionosphere beyond satellite altitudes, followed by subsequent plasma diffusion to higher latitudes along magnetic field lines, results in the formation of trough‐like plasma depletions around the magnetic equator and density enhancement at higher latitudes. The detection of plasma bubbles in the Asian sector during the non‐bubble season (January) is likely associated with the uplift of the ionosphere at the sunset terminator.

Extreme (>20 nT/s) geomagnetic disturbances (GMDs, also denoted as MPEs—magnetic perturbation events)—impulsive nighttime disturbances with time scale ∼5–10 min, have sufficient amplitude to cause bursts of geomagnetically induced currents (GICs) that can damage technical infrastructure. In this study, we present occurrence statistics for extreme GMD events from five stations in the MACCS and AUTUMNX magnetometer arrays in Arctic Canada at magnetic latitudes ranging from 65° to 75°. We report all large (≥6 nT/s) and extreme GMDs from these stations from 2011 through 2022 to analyze variations of GMD activity over a full solar cycle and compare them to those found in three earlier studies. GMD activity between 2011 and 2022 did not closely follow the sunspot cycle, but instead was lowest during its rising phase and maximum (2011–2014) and highest during the early declining phase (2015–2017). Most of these GMDs, especially the most extreme, were associated with high‐speed solar wind streams (Vsw >600 km/s) and steady solar wind pressure. All extreme GMDs occurred within 80 min after substorm onsets, but few within 5 min. Multistation data often revealed a poleward progression of GMDs, consistent with a tailward retreat of the magnetotail reconnection region. These observations indicate that extreme GIC hazard conditions can occur for a variety of solar wind drivers and geomagnetic conditions, not only for fast‐coronal mass ejection driven storms.

Inter-hemispheric asymmetry (IHA) in Earth’s ionosphere–thermosphere (IT) system can be associated with high-latitude forcing that intensifies during storm time, e.g., ion convection, auroral electron precipitation, and energy deposition, but a comprehensive understanding of the pathways that generate IHA in the IT is lacking. Numerical simulations can help address this issue, but accurate specification of high-latitude forcing is needed. In this study, we utilize the Active Magnetosphere and Planetary Electrodynamics Response Experiment-revised fieldaligned currents (FACs) to specify the high-latitude electric potential in the Global Ionosphere and Thermosphere Model (GITM) during the October 8–9, 2012, storm. Our result illustrates the advantages of the FAC-driven technique in capturing high-latitude ion drift, ion convection equatorial boundary, and the storm-time neutral density response observed by satellite. First, it is found that the cross-polar-cap potential, hemispheric power, and ion convection distribution can be highly asymmetric between two hemispheres with a clear B_ydependence in the convection equatorial boundary. Comparison with simulation based on mirror precipitation suggests that the convection distribution is more sensitive to FAC, while its intensity also depends on the ionospheric conductance-related precipitation. Second, the IHA in the neutral density response closely follows the IHA in the total Joule heating dissipation with a time delay. Stronger Joule heating deposited associated with greater high-latitude electric potential in the southern hemisphere during the focus period generates more neutral density as well, which provides some evidences that the high-latitude forcing could become the dominant factor to IHAs in the thermosphere when near the equinox. Our study improves the understanding of storm-time IHA in high-latitude forcing and the IT system.

Understanding the contributions of transcription factor DNA binding sites to transcriptional enhancers is a significant challenge. We developed Quantitative enhancer-FACS-Seq for highly parallel quantification of enhancer activities from a genomically integrated reporter inDrosophila melanogasterembryos. We investigate the contributions of the DNA binding motifs of four poorly characterized TFs to the activities of twelve embryonic mesodermal enhancers. We measure quantitative changes in enhancer activity and discover a range of epistatic interactions among the motifs, both synergistic and alleviating. We find that understanding the regulatory consequences of TF binding motifs requires that they be investigated in combination across enhancer contexts.