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Abstract We report a detailed analysis of GAMA 376183, a powerful, heavily obscured active galactic nucleus (AGN) hosted by a low-mass galaxy (M_⋆ ≈ 10¹⁰M_⊙) likely experiencing a galaxy merger. The source was initially identified due to its remarkably strong [Nev]λ3426 emission, exhibiting a rest-frame equivalent width of ≈48 Å. We present ∼100 ks Nuclear Spectroscopic Telescope Array (or NuSTAR) follow-up observations, confirming its heavily obscured nature with a column density (in cm⁻²) of $\log N_{\mathrm{H}}=23.3_{-1.2}^{+0.4}$ and an intrinsic 2–10 keV luminosity (in erg s⁻¹) of $\log L_{\mathrm{X},\mathrm{int}}=42.92_{-0.20}^{+0.24}$ . GAMA 376183 thus represents one of the few known heavily obscured AGNs in low-mass galaxies. Its estimated Eddington ratio isλ_Edd ≈ 0.8, indicative of rapid black hole growth. High-resolution optical images reveal a disturbed, likely merging morphology, while its multiwavelength spectral energy distribution indicates a recent starburst in its host galaxy. These pieces of evidence suggest that the ongoing merger has triggered both the heavily obscured, Eddington-limited accretion and the starburst, making GAMA 376183 a rare observed case in low-mass galaxies. Overall, this unique source demonstrates that (i) [Nev] can help identify heavily obscured low-mass AGNs, and (ii) the merger-driven coevolution framework established for massive galaxies may also extend to low-mass galaxies. 

Abstract Dust-obscured galaxies (DOGs) with extremely red optical-to-infrared colors are often associated with intense starburst and active galactic nucleus (AGN) activity. Studying DOGs can provide insights into the processes that drive the growth of galaxies and their central supermassive black holes. However, the general DOG population is heterogeneous, spanning a wide range of evolutionary stages, and has X-ray obscuring column densities (N_H) covering low to high levels. In this work, we focus on seven high Eddington ratio DOGs ( $\log {\lambda }_{\mathrm{Edd}}\gtrsim -0.5$ ) to examine their X-ray obscuration properties using new and archival X-ray observations. We confirm that these systems are generally heavily obscured, with six out of seven havingN_H ≳ 10²³cm⁻²and three out of seven havingN_H ≳ 10²⁴cm⁻². Based on the observed similarity with the rare Hot DOG population, we argue that both high-λ_EddDOGs and Hot DOGs likely trace the postmerger phase, during which AGNs are enshrouded by large columns of dust-rich material. 

Abstract It is well established that cosmic supermassive black hole (SMBH) growth peaks atz ≈ 1.5–2, followed by a strong decline of ≈1–1.5 dex toward the present day, with the comoving number density of higher-luminosity active galactic nuclei (AGNs) peaking at higher redshift (referred to as “AGN downsizing”). We leverage the best current measurements of the SMBH accretion distribution, based upon data from nine well-characterized extragalactic fields with a “wedding-cake” design, to investigate and quantify the drivers of the drastic decline in cosmic SMBH growth. The decline in the typical Eddington ratio (λ_Edd) of AGNs (decreasing by ≈1.35 dex fromz ≈ 1.5–2 toz ≈ 0.2) is the dominant driver for the broad decline in SMBH growth, rather than a shift of accretion activity to less-massive SMBHs. Asλ_Edddecreases toward lower redshift, the primary contributor to the cosmic SMBH accretion density (ρ_BHAR) has shifted from high-λ_EddAGNs to low-λ_EddAGNs, even though the latter always dominate the comoving AGN number density atz < 4. We also find that the decline in SMBH growth toward lower SMBH mass in less-massive galaxies is primarily due to the decreasing outburst luminosity rather than the duty cycle. 

Abstract The 3D genome organization plays a key role in regulating interactions among chromosomal loci. While Chromosome Conformation Capture (3C)-based methods have provided static snapshots of chromatin architecture, the kinetics of chromosomal encounters in live cells remain poorly characterized. In this study, we employ Chemically Induced Chromosomal Interaction (CICI) to measure encounter times between multiple loci pairs in G1-arrested budding yeast. Our results show that chromosome motion closely follows the Rouse polymer model, with similar diffusion parameters at all tested loci. Surprisingly, we find that long-range intra-chromosomal encounters occur significantly faster than inter-chromosomal encounters at similar 3D distances. Using targeted depletion experiments, we identify condensin, but not cohesin, as the complex responsible for these rapid intra-chromosomal interactions. This is further supported by Hi-C analysis, which reveals that condensin promotes long-distance intra-chromosomal interactions in G1 yeast. Through polymer simulations, we estimate that condensin extrudes chromatin at ∼2 kb/s with a density of one complex per 1-2 Mb and a processivity of 120-220 kb. These findings uncover a novel role for condensin in shaping the interphase genome organization and provide new insights into chromosomal search dynamicsin vivo. 

Abstract Dust-obscured galaxies (DOGs) containing central supermassive black holes (SMBHs) that are rapidly accreting (i.e., having high Eddington ratios,λ_Edd) may represent a key phase closest to the peak of both the black hole and galaxy growth in the coevolution framework for SMBHs and galaxies. In this work, we present a 68 ks XMM-Newton observation of the high-λ_EddDOG J1324+4501 atz∼ 0.8, which was initially observed by Chandra. We analyze the XMM-Newton spectra jointly with archival Chandra spectra. In performing a detailed X-ray spectral analysis, we find that the source is intrinsically X-ray luminous with $\log (L_{\mathrm{X}}$ /erg ${\mathrm{s}}^{-1})={44.71}_{-0.12}^{+0.08}$ and heavily obscured with $\log \left(N_{\mathrm{H}}/{\mathrm{cm}}^{-2}\right)={23.43}_{-0.13}^{+0.09}$ . We further utilize UV-to-IR archival photometry to measure and fit the source’s spectral energy distribution to estimate its host-galaxy properties. We present a supplementary comparison sample of 21 X-ray luminous DOGs from the XMM-SERVS survey with sufficient (>200) 0.5–10 keV counts to perform a similarly detailed X-ray spectral analysis. Of the X-ray luminous DOGs in our sample, we find that J1324+4501 is the most remarkable, possessing one of the highest X-ray luminosities, column densities, and star formation rates. We demonstrate that J1324+4501 is in an extreme evolutionary stage where SMBH accretion and galaxy growth are at their peaks. 

Abstract The coevolution of supermassive black holes and their host galaxies represents a fundamental question in astrophysics. One approach to investigating this question involves comparing the star formation rates (SFRs) of active galactic nuclei (AGNs) with those of typical star-forming galaxies. At relatively low redshifts (z≲ 1), radio AGNs manifest diminished SFRs, indicating suppressed star formation, but their behavior at higher redshifts is unclear. To examine this, we leveraged galaxy and radio-AGN data from the well-characterized W-CDF-S, ELAIS-S1, and XMM-LSS fields. We established two mass-complete reference star-forming galaxy samples and two radio-AGN samples, consisting of 1763 and 6766 radio AGNs, the former being higher in purity and the latter more complete. We subsequently computed star-forming fractions (f_SF; the fraction of star-forming galaxies to all galaxies) for galaxies and radio-AGN host galaxies and conducted a robust comparison between them up toz≈ 3. We found that the tendency for radio AGNs to reside in massive galaxies primarily accounts for their lowf_SF, which also shows a strong negative dependence uponM_⋆and a strong positive evolution withz. To investigate further the star formation characteristics of those star-forming radio AGNs, we constructed the star-forming main sequence (MS) and investigated the behavior of the position of AGNs relative to the MS atz≈ 0–3. Our results reveal that radio AGNs display lower SFRs than star-forming galaxies in the low-zand high-M_⋆regime and, conversely, exhibit comparable or higher SFRs than MS star-forming galaxies at higher redshifts or lowerM_⋆. 

Abstract Supermassive black holes (SMBHs) can grow through both accretion and mergers. It is still unclear how SMBHs evolve under these two channels from high redshifts to the SMBH population we observe in the local Universe. Observations can directly constrain the accretion channel but cannot effectively constrain mergers yet, while cosmological simulations provide galaxy merger information but can hardly return accretion properties consistent with observations. In this work, we combine the observed accretion channel and the simulated merger channel, taking advantage of observations and cosmological simulations, to depict a realistic evolution pattern of the SMBH population. With this methodology, we can derive the scaling relation between the black hole mass (M_BH) and host-galaxy stellar mass (M_⋆), and the local black hole mass function (BHMF). Our scaling relation is lower than those based on dynamically measuredM_BH, supporting the claim that dynamically measured SMBH samples may be biased. We show that the scaling relation has little redshift evolution. The BHMF steadily increases fromz= 4 toz= 1 and remains largely unchanged fromz= 1 toz= 0. The overall SMBH growth is generally dominated by the accretion channel, with possible exceptions at high mass (M_BH≳ 10⁸M_⊙orM_⋆≳ 10¹¹M_⊙) and low redshift (z≲ 1). We also predict that around 25% of the total SMBH mass budget in the local Universe may be locked within long-lived, wandering SMBHs, and the wandering mass fraction and wandering SMBH counts increase withM_⋆. 

Abstract Dust-obscured galaxies (DOGs) are enshrouded by dust and many are believed to host accreting supermassive black holes (SMBHs), which makes them unique objects for probing the coevolution of galaxies and SMBHs. We select and characterize DOGs in the 13 deg²XMM-Spitzer Extragalactic Representative Volume Survey (XMM-SERVS), leveraging the superb multiwavelength data—from X-rays to radio. We select 3738 DOGs atz≈ 1.6–2.1 in XMM-SERVS, while maintaining good data quality without introducing significant bias. This represents the largest DOG sample with thorough multiwavelength source characterization. Spectral energy distribution modeling shows DOGs are a heterogeneous population consisting of both normal galaxies and active galactic nuclei (AGNs). Our DOGs are massive ( $\log M_{\star }/M_{☉}\approx 10.7-11.3$ ), 174 are detected in X-rays, and they are generally radio-quiet systems. X-ray detected DOGs are luminous and are moderately to heavily obscured in X-rays. Stacking analyses for the X-ray undetected DOGs show highly significant average detections. Critically, we compare DOGs with matched galaxy populations. DOGs have similar AGN fractions compared with typical galaxy populations. X-ray detected DOGs have higherM_⋆and higher X-ray obscuration, but they are not more star-forming than typical X-ray AGNs. Our results potentially challenge the relevance of the merger-driven galaxy-SMBH coevolution framework for X-ray detected DOGs. 

Some transcription factors (TFs) can form liquid-liquid phase separated (LLPS) condensates. However, the functions of these TF condensates in 3D genome organization and gene regulation remain elusive. In response to methionine (met) starvation, budding yeast TF Met4 and a few co-activators, including Met32, induce a set of genes involved in met biosynthesis. Here, we show that the endogenous Met4 and Met32 form co-localized puncta-like structures in yeast nuclei upon met depletion. Recombinant Met4 and Met32 form mixed droplets with LLPS properties in vitro. In relation to chromatin, Met4 puncta co-localize with target genes, and at least a subset of these target genes are clustered in 3D in a Met4-dependent manner. A MET3pr-GFP reporter inserted near several native Met4 binding sites becomes co-localized with Met4 puncta and displays enhanced transcriptional activity. A Met4 variant with a partial truncation of an intrinsically disordered region (IDR) shows less puncta formation, and this mutant selectively reduces the reporter activity near Met4 binding sites to the basal level. Overall, these results support a model where Met4 and co-activators form condensates to bring multiple target genes into a vicinity with higher local TF concentrations, which facilitates a strong response to methionine depletion. 

Abstract The growth of supermassive black holes is strongly linked to their galaxies. It has been shown that the population mean black hole accretion rate ( $\overline{\mathrm{BHAR}}$ ) primarily correlates with the galaxy stellar mass (M_⋆) and redshift for the general galaxy population. This work aims to provide the best measurements of $\overline{\mathrm{BHAR}}$ as a function ofM_⋆and redshift over ranges of 10^9.5<M_⋆< 10¹²M_⊙andz< 4. We compile an unprecedentedly large sample with 8000 active galactic nuclei (AGNs) and 1.3 million normal galaxies from nine high-quality survey fields following a wedding cake design. We further develop a semiparametric Bayesian method that can reasonably estimate $\overline{\mathrm{BHAR}}$ and the corresponding uncertainties, even for sparsely populated regions in the parameter space. $\overline{\mathrm{BHAR}}$ is constrained by X-ray surveys sampling the AGN accretion power and UV-to-infrared multiwavelength surveys sampling the galaxy population. Our results can independently predict the X-ray luminosity function (XLF) from the galaxy stellar mass function (SMF), and the prediction is consistent with the observed XLF. We also try adding external constraints from the observed SMF and XLF. We further measure $\overline{\mathrm{BHAR}}$ for star-forming and quiescent galaxies and show that star-forming $\overline{\mathrm{BHAR}}$ is generally larger than or at least comparable to the quiescent $\overline{\mathrm{BHAR}}$ .