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  1. Free, publicly-accessible full text available May 1, 2026
  2. Free, publicly-accessible full text available January 1, 2026
  3. A long-term record of global mean surface temperature (GMST) provides critical insight into the dynamical limits of Earth’s climate and the complex feedbacks between temperature and the broader Earth system. Here, we present PhanDA, a reconstruction of GMST over the past 485 million years, generated by statistically integrating proxy data with climate model simulations. PhanDA exhibits a large range of GMST, spanning 11° to 36°C. Partitioning the reconstruction into climate states indicates that more time was spent in warmer rather than colder climates and reveals consistent latitudinal temperature gradients within each state. There is a strong correlation between atmospheric carbon dioxide (CO2) concentrations and GMST, identifying CO2as the dominant control on variations in Phanerozoic global climate and suggesting an apparent Earth system sensitivity of ~8°C. 
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  4. Paleo-CO2 reconstructions are integral to understanding the evolution of Earth system processes and their interactions given that atmospheric-CO2 concentrations are intrinsically linked to planetary function. In this talk, we use several case studies, spanning the 3 Phanerozoic Eras, to illustrate the potential of paleo-CO2 records to constrain the magnitude and state-dependency of equilibrium climate sensitivity, to advance our understanding of global biogeochemical cycles, to test the sensitivity of Earth System modeled atmospheric and oceanic circulation to PCO2 over a range of climate states, and to interrogate ecosystem—CO2—climate linkages and physiological responses to CO2. Further advances in these areas, however, are dependent on how well we ‘know’ paleo-CO2 estimates. CO2 estimates exist for much of the past half-billion years, but the degree to which the accuracy and precision of these estimates are constrained is quite variable, leading to substantial uncertainty and inconsistency in paleo-CO2 estimates. Potential sources of this uncertainty and inconsistency include an incomplete understanding of how environmental and ecophysiological conditions and processes imprint the CO2 proxy signals we measure, of the sensitivity of the CO2 estimates to this uncertainty, and differences in approaches to assigning uncertainties to CO2 estimates, among other factors. Application of newly established screening criteria, defined as part of an effort to improve our understanding of how atmospheric CO2 has varied through the Cenozoic, illustrates how the majority of pre-Cenozoic estimates are unreliable in their current form. To address these issues and to advance paleo-CO2 reconstruction, we introduce CO2PIP, a new community-scale project that takes a two-step approach to building the next generation Phanerozoic-CO2 record. Collective efforts are modernizing existing terrestrial-based CO2 estimates through additional analyses, measurements and proxy system modeling to constrain critical parameters used to estimate paleo-CO2. A set of forward proxy system models being developed in collaboration with the CO2 community, will provide a quantified representation of proxy sensitivities to environmental and ecophysiological conditions and processes that govern the CO2 signals. Ultimately, statistical inversion analysis of the simulated and modernized proxy datasets will be used to revise individual CO2 records and to build a new integrated model-data-constrained CO2 record for the Phanerozoic. 
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