The Sahel is highly sensitive to flooding, droughts, and wildfires, risking food and other resources on which nearly 100 million people depend. Understanding how natural variations of precipitation and vegetation fluctuate during high-amplitude glacial- interglacial cycles can help constrain the regional sensitivity to a wide range of external forcings. Further, the interactions between climate and ecosystem changes remain uncertain for sub-Saharan Africa due to the lack of long, highly-resolved, quantitative, terrestrial records. Here we present precipitation and vegetation records from ~215 ka to present, derived from long leaf wax hydrogen (δDwax) and carbon (δ13Cwax) isotopes, respectively. These geochemical records are derived from ODP Site 959 in the Gulf of Guinea, where westerly winds and major river systems transport Western Sahel-sourced terrestrial leaf waxes. We find that, unlike many African records that are precessionally- driven, obliquity plays an important role in West African late Pleistocene hydroclimate, suggesting that a cross-equatorial insolation gradient may be more important in this area and certainly that drivers of orbital-scale precipitation change are regionally-specific. Further, vegetation changes appear to have a complex relationship with hydroclimate over this mid-late Pleistocene interval. A potential shift in this climate-environment coupling at MIS6 ~130 ka, which is a time when there is also a shift in forcing mechanisms in East Africa, suggests that the global boundary condition changes associated with large glacial- interglacial cycling may affect equatorial climate.
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Late Middle Pleistocene Elephants from Natodomeri, Kenya and the Disappearance of Elephas (Proboscidea, Mammalia) in Africa
Comparative morphometric study of recently recovered fossil elephant molars from Natodomeri, Kenya identifies them as belonging to Elephas jolensis and confirms the presence of this species in Members I and II of the Kibish Formation. Improved datation of these geological units constrains them between 205 and 130 ka. Elephas jolensis is also reported from localities in northern, northwestern, eastern, and southern Africa. Thus, including its Natodomeri occurrence, E. jolensis appears to have been pan- African in distribution. Despite the wide geographic distribution of the species, molars of E. jolensis are remarkably uniform morphometrically. They are characterized by their extreme hypsodonty, high amplitude of enamel folding, high lamellar frequency, and plates that are anteroposteriorly thick relative to transverse valley interval spacing. In addition, they exhibit only a modest number of plates (<20 in M3/m3). Elephas jolensis either evolved from or represents the last stage of Elephas recki, the dominant elephant species in East Africa during the late Pliocene-Pleistocene. The dental morphology and isotopic composition of E. jolensis indicates that, like E. recki, it was a dedicated grazer. In the Kibish Formation, E. jolensis is succeeded by Loxodonta africana at 130 ka, coincident with an intensely cool, dry interval marked by episodes of extreme drought. This marked the extirpation of Elephas on the continent. The intensity and increased rate of climate fluctuation may have played an important role in the demise of the specialist, grazing E. recki-E. jolensis lineage in favor of a generalist, mixed feeder such as L. africana. Keywords Natodomeri, Kenya . Kibish Formation . Elephantidae . Elephas jolensis . Late middle Pleistocene
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- Award ID(s):
- 1740383
- PAR ID:
- 10125003
- Date Published:
- Journal Name:
- Journal of Mammalian Evolution
- ISSN:
- 1064-7554
- Format(s):
- Medium: X
- Sponsoring Org:
- National Science Foundation
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