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Abstract PremiseDioecy (separate sexes) has independently evolved numerous times across the angiosperm phylogeny and is recently derived in many lineages. However, our understanding is limited regarding the evolutionary mechanisms that drive the origins of dioecy in plants. The recent and repeated evolution of dioecy across angiosperms offers an opportunity to make strong inferences about the ecological, developmental, and molecular factors influencing the evolution of dioecy, and thus sex chromosomes. The genusAsparagus(Asparagaceae) is an emerging model taxon for studying dioecy and sex chromosome evolution, yet estimates for the age and origin of dioecy in the genus are lacking. MethodsWe use plastome sequences and fossil time calibrations in phylogenetic analyses to investigate the age and origin of dioecy in the genusAsparagus. We also review the diversity of sexual systems present across the genus to address contradicting reports in the literature. ResultsWe estimate that dioecy evolved once or twice approximately 2.78−3.78 million years ago inAsparagus, of which roughly 27% of the species are dioecious and the remaining are hermaphroditic with monoclinous flowers. ConclusionsOur findings support previous work implicating a young age and the possibility of two origins of dioecy inAsparagus, which appear to be associated with rapid radiations and range expansion out of Africa. Lastly, we speculate that paleoclimatic oscillations throughout northern Africa may have helped set the stage for the origin(s) of dioecy inAsparagusapproximately 2.78−3.78 million years ago. 

Abstract. The role of clouds in the Arctic radiation budget is not well understood. Ground-based and airborne measurements provide valuable data to test and improve our understanding. However, the ground-based measurements are intrinsically sparse, and the airborne observations are snapshots in time and space. Passive remote sensing measurements from satellite sensors offer high spatial coverage and an evolving time series, having lengths potentially of decades. However, detecting clouds by passive satellite remote sensing sensors is challenging over the Arctic because of the brightness of snow and ice in the ultraviolet and visible spectral regions and because of the small brightness temperature contrast to the surface. Consequently, the quality of the resulting cloud data products needs to be assessed quantitatively. In this study, we validate the cloud data products retrieved from the Advanced Very High Resolution Radiometer (AVHRR) post meridiem (PM) data from the polar-orbiting NOAA-19 satellite and compare them with those derived from the ground-based instruments during the sunlit months. The AVHRR cloud data products by the European Space Agency (ESA) Cloud Climate Change Initiative (Cloud_CCI) project uses the observations in the visible and IR bands to determine cloud properties. The ground-based measurements from four high-latitude sites have been selected for this investigation: Hyytiälä (61.84∘ N, 24.29∘ E), North Slope of Alaska (NSA; 71.32∘ N, 156.61∘ W), Ny-Ålesund (Ny-Å; 78.92∘ N, 11.93∘ E), and Summit (72.59∘ N, 38.42∘ W). The liquid water path (LWP) ground-based data are retrieved from microwave radiometers, while the cloud top height (CTH) has been determined from the integrated lidar–radar measurements. The quality of the satellite products, cloud mask and cloud optical depth (COD), has been assessed using data from NSA, whereas LWP and CTH have been investigated over Hyytiälä, NSA, Ny-Å, and Summit. The Cloud_CCI COD results for liquid water clouds are in better agreement with the NSA radiometer data than those for ice clouds. For liquid water clouds, the Cloud_CCI COD is underestimated roughly by 3 optical depth (OD) units. When ice clouds are included, the underestimation increases to about 5 OD units. The Cloud_CCI LWP is overestimated over Hyytiälä by ≈7 g m−2, over NSA by ≈16 g m−2, and over Ny-Å by ≈24 g m−2. Over Summit, CCI LWP is overestimated for values ≤20 g m−2 and underestimated for values >20 g m−2. Overall the results of the CCI LWP retrievals are within the ground-based instrument uncertainties. To understand the effects of multi-layer clouds on the CTH retrievals, the statistics are compared between the single-layer clouds and all types (single-layer + multi-layer). For CTH retrievals, the Cloud_CCI product overestimates the CTH for single-layer clouds. When the multi-layer clouds are included (i.e., all types), the observed CTH overestimation becomes an underestimation of about 360–420 m. The CTH results over Summit station showed the highest biases compared to the other three sites. To understand the scale-dependent differences between the satellite and ground-based data, the Bland–Altman method is applied. This method does not identify any scale-dependent differences for all the selected cloud parameters except for the retrievals over the Summit station. In summary, the Cloud_CCI cloud data products investigated agree reasonably well with those retrieved from ground-based measurements made at the four high-latitude sites.