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Abstract The Campanian Two Medicine Formation of northwestern Montana, USA, is richly fossiliferous, and discoveries made within the unit over the past century have greatly advanced our appreciation of dinosaur paleobiology and evolution. Previously undifferentiated from a lithostratigraphic perspective, the formation is now subdivided into four new members that include (from base to top) (1) the Rock City Member, (2) the Shields Crossing Member, (3) the Hagans Crossing Member, and (4) the Flag Butte Member. These new formal units and their associated fossil occurrences are also now included in an age model founded on eight high-resolution chemical abrasion–isotope dilution–thermal ionization mass spectrometry (CA-ID-TIMS) U-Pb ages. New age data confirm that the Two Medicine Formation accumulated during much of the Campanian, with deposition spanning ca. 82.4 Ma to 74.4 Ma. New age data further indicate that a major reorganization of depositional systems, marked by a shift from predominantly lacustrine to alluvial facies and accompanied by a dramatic increase in accommodation, transpired near the base of the new Flag Butte Member at ca. 76.3 Ma. This change in depositional regime correlates in age with the Judith River–Belly River discontinuity, which marks the contact between the McClelland Ferry and Coal Ridge Members in the Judith River Formation and coincides with the onset of the Bearpaw transgression in north-central Montana. The new lithostratigraphic and chronostratigraphic framework for the Two Medicine Formation serves to contextualize and calibrate the formation’s rich dinosaur fossil record, which can now be interrogated with increased clarity and precision. These results also provide ground truth for numerical models that explore the structure of the fossil record in relation to alluvial architecture and terrestrial sequence stratigraphy. 

Taphonomic processes create bias in the fossil record, and understanding these processes is integral to interpreting the record of extinct life worldwide. Bones preserved in fluvial environments make up a substantial part of the vertebrate fossil record. These bones have often been transported varying distances from the location of death before becoming buried. Experiments in flumes and natural settings have explored the fluvial taphonomy of mammal skeletons, but the taphonomy of other terrestrial vertebrates, especially extinct clades, has only been sparingly studied directly. Hadrosauroids are a dinosaur clade known from extensive remain throughout the Cretaceous and across the globe, making them an ideal group for taphonomic study. Previous examinations regarding the fluvial taphonomy of their skeletons have often applied bone transport groups derived from classic studies on mammals. Some researchers have raised concerns that the morphologies of non-mammalian bones would not exhibit the same hydraulic properties as mammals, producing different transport patterns. Here, we investigate hadrosauroid bone transport under various flow conditions through actualistic flume experiments using 3d printed models with comparable densities to real bone. We aimed to characterize the timing of transport of different elements (Voorhies Groups), orientation of bones relative to flow direction, and bone surface abrasion patterns. Some elements behave similarly to those described in mammals. As would be expected from previous work, relatively heavy bones such as the femur tend to move last, acting as lag elements. Lighter elements such as the scapula and radius tended to begin moving at much lower flow speeds. Because dinosaur pelvic bones are not fused as in mammals, we observed that the isolated pubis is often among the first elements to commence movement, often rotating or sliding along the bed. Cylindrical limb bones tend to roll or slide along the bed, orienting to be parallel to flow faster or slower depending on element size and flow velocity. Bones with more complex shapes, such as the curved and concave blade of the scapula, moved in less straightforward and unique ways, even vaulting over other bones. We also found that burial by fine silt and mud could be achieved relatively quickly even at slower flow speeds, and burial by sand played an important part in inhibiting transport in higher flow regimes. 

Terrestrial strata of the Judith River−Belly River wedge, widely exposed in the plains of north-central Montana, southern Alberta, and southwestern Saskatchewan, were pivotal in early stratigraphic investigations of the Western Interior of North America and are renowned to this day for their spectacular preservation of Late Cretaceous fossils, most notably dinosaurs. Correlation of the Judith River Formation in Montana with the Foremost, Oldman, and Dinosaur Park Formations (= Belly River Group) in Canada has been challenging for a variety of reasons, including lithostratigraphic complexities, legacy bentonite ages of limited comparability, and distinctly different stratigraphic models on opposite sides of the international border. An updated model calibrated with U-Pb zircon ages provides an improved framework for stratigraphic analysis. New geochronology indicates that the Oldman−Dinosaur Park discontinuity in Dinosaur Provincial Park correlates in age with the mid-Judith discontinuity in the Judith River Formation in Montana, which is interpreted as an expansion surface linked to a major pulse of accommodation and onset of the Bearpaw transgression at ca. 76.3 Ma. The regionally expressed shift in alluvial facies marking the mid-Judith discontinuity can be traced in well logs from Montana to southern Canada, where it loses distinction and transitions to a subsurface signature typical of the Oldman−Dinosaur Park discontinuity, which in turn can be traced north to Dinosaur Provincial Park and beyond. Across this expanse, both discontinuities parallel the Eagle/Milk River shoulder at approximately the same stratigraphic height, confirming their chronostratigraphic significance. These findings have clear implications for regional correlation and the evolution of alluvial depositional systems in a foreland basin setting, and they afford an opportunity to evaluate existing interpretations and advance understanding of the stratigraphy and paleontology of the Judith River−Belly River wedge. The term “Judith River−Belly River discontinuity” should be used henceforth to refer to the chronostratigraphically significant stratal discontinuity that subdivides the Judith River−Belly River wedge throughout the plains of north-central Montana, southern Alberta, and southwestern Saskatchewan. 

The fossil record of mammaliaforms (mammals and their closest relatives) of the Mesozoic era from the southern supercontinent Gondwana is far less extensive than that from its northern counterpart, Laurasia. Among Mesozoic mammaliaforms, Gondwanatheria is one of the most poorly known clades, previously represented by only a single cranium and isolated jaws and teeth. As a result, the anatomy, palaeobiology and phylogenetic relationships of gondwanatherians remain unclear. Here we report the discovery of an articulated and very well-preserved skeleton of a gondwanatherian of the latest age (72.1–66 million years ago) of the Cretaceous period from Madagascar that we assign to a new genus and species, Adalatherium hui. To our knowledge, the specimen is the most complete skeleton of a Gondwanan Mesozoic mammaliaform that has been found, and includes the only postcranial material and ascending ramus of the dentary known for any gondwanatherian. A phylogenetic analysis including the new taxon recovers Gondwanatheria as the sister group to Multituberculata. The skeleton, which represents one of the largest of the Gondwanan Mesozoic mammaliaforms, is particularly notable for exhibiting many unique features in combination with features that are convergent on those of therian mammals. This uniqueness is consistent with a lineage history for A. hui of isolation on Madagascar for more than 20 million years.