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Abstract There is a consensus that volcanism along the East African Rift System (EARS) is related to plume activities. However, because of our limited knowledge of the local lithospheric mantle, the dynamics of the plume are poorly constrained by magma chemistry. The Turkana Basin is one of the best places to study plume‐related volcanism because the lithospheric mantle there is unusually thin. New Ar‐Ar geochronology and geochemical data on lavas from western Turkana show that Eocene volcanics have relatively low²⁰⁶Pb/²⁰⁴Pb (<19.1) and high εNd (>3.78). Their relatively high Ba/Rb (35–78) ratios suggest contributions from the shallow lithospheric mantle. Oligo‐Miocene Turkana volcanics have HIMU‐ and EMI‐ type enriched mantle signatures with overall lower Ba/Rb ratios, which is consistent with partial melting of plume material. Pliocene and younger Turkana volcanics have low Ba/Rb and Sr‐Nd‐Pb isotope ratios that resemble those of Ethiopian volcanics with elevated³He/⁴He ratios. This temporal variation can be reconciled with a layered plume model where an outer layer of ancient recycled oceanic crust and sediment overlies more primitive lower mantle material. Beneath Ethiopia, the outer layer of the plume is either missing or punctured by the delamination of the thicker overlying lithospheric mantle atca.30 Ma, an event that would have facilitated the rapid upwelling of the inner portion of the plume and triggered the Ethiopian flood volcanism. The outer layer of the plume may be thicker in the southern EARS, which could explain the occurrence of young HIMU‐ and EMI‐type volcanics with primordial noble gas signatures. 

Variability in resource availability is hypothesized to be a significant driver of primate adaptation and evolution, but most paleoclimate proxies cannot recover environmental seasonality on the scale of an individual lifespan. Oxygen isotope compositions (δ 18 O values) sampled at high spatial resolution in the dentitions of modern African primates ( n = 2,352 near weekly measurements from 26 teeth) track concurrent seasonal precipitation, regional climatic patterns, discrete meteorological events, and niche partitioning. We leverage these data to contextualize the first δ 18 O values of two 17 Ma Afropithecus turkanensis individuals from Kalodirr, Kenya, from which we infer variably bimodal wet seasons, supported by rainfall reconstructions in a global Earth system model. Afropithecus ’ δ 18 O fluctuations are intermediate in magnitude between those measured at high resolution in baboons ( Papio spp.) living across a gradient of aridity and modern forest-dwelling chimpanzees ( Pan troglodytes verus ). This large-bodied Miocene ape consumed seasonally variable food and water sources enriched in 18 O compared to contemporaneous terrestrial fauna ( n = 66 fossil specimens). Reliance on fallback foods during documented dry seasons potentially contributed to novel dental features long considered adaptations to hard-object feeding. Developmentally informed microsampling recovers greater ecological complexity than conventional isotope sampling; the two Miocene apes ( n = 248 near weekly measurements) evince as great a range of seasonal δ 18 O variation as more time-averaged bulk measurements from 101 eastern African Plio-Pleistocene hominins and 42 papionins spanning 4 million y. These results reveal unprecedented environmental histories in primate teeth and suggest a framework for evaluating climate change and primate paleoecology throughout the Cenozoic.