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Abstract We evaluate five commonly‐applied criteria to validate that a climate model is in so‐called “quasi‐equilibrium,” using a suite of five simulations with CO₂concentrations between 1× and 16× Pre‐Industrial values. We find that major changes in ocean circulation can occur after common thermal equilibrium criteria are reached, such as a small Top of Atmosphere radiative flux imbalance, or weak trends in surface air temperature, sea surface temperature, and deep ocean temperature. Ocean circulation change, in turn, impact high‐latitude SAT, sea ice, and the Inter‐tropical Convergence Zone position. For future modeling studies and intercomparison projects aiming for an ocean in quasi‐equilibrium, we suggest that time series of key meridional overturning circulation (MOC) metrics in the Atlantic, Pacific, and Southern Ocean are saved, and that MOC trends are less than 1 Sv/1000 years, and DOT trends less than 0.1°C/century for the final 1000 years of the simulations. 

Abstract The Miocene (∼23–5 Ma) experienced substantial paleogeographic changes, including the shoaling of the Panama Seaway and closure of the Tethys Seaway, which altered exchange pathways between the Pacific and Atlantic Oceans. Changes in continental configuration and topography likely also influenced global wind patterns. Here, we investigate how these changes affected surface wind‐driven gyre circulation and interbasin volume transport using 14 fully coupled climate model simulations of the early and middle Miocene. The North and South Atlantic gyres, along with the South Pacific gyre, are weaker in the Miocene simulations compared to pre‐industrial (PI), while the North Pacific gyres are stronger. These changes largely follow the wind stress curl and basin width changes. Westward flow through the Panama Seaway occurs only in early Miocene simulations when the Tethys Seaway is open and transports are strongly westward. As the Tethys transport declines, flow across the Panama Seaway gradually reverses from westward (into the Pacific) to eastward (into the Atlantic). In simulations with a closed Tethys Seaway, the Panama transport is consistently eastward. The Southern Hemisphere westerlies are weaker than PI in all simulations, contributing to a reduced Antarctic Circumpolar Current (ACC) in 11 of the 14 cases. In the remaining three, a stronger ACC is simulated, likely due to a combination of enhanced meridional density gradients and model‐dependent sensitivities. These findings highlight how changes in Miocene seaways and wind patterns reshaped ocean circulation, influencing interbasin exchange, thermohaline properties, and global climate. 

Abstract The tropical Pacific climate has an outsized impact on global climate, yet future projections are poorly constrained. Data‐model comparisons from the mid‐Pliocene warm period (3.3 million years ago) can help investigate warm climate dynamics and evaluate model behavior. Here we compare proxy records to PlioMIP2 models and a model with modified cloud albedo. Relative to modern, the mid‐Pliocene warm period records show subsurface warming across the tropical Pacific, strong eastern Pacific surface warming and weak western Pacific surface warming. Using clustering analyses to group model behavior relative to the proxy data, we find the model cluster with the best fit with the proxy data has enhanced warming in mid‐latitude thermocline source water regions which connect to the equator through the ventilated thermocline. Our study shows tropical ocean heat content during the mid‐Pliocene warm period was higher than today and has broad implications for the ocean's ability to absorb anthropogenic heat. 

Abstract The Miocene (∼23–5 Ma) is a past warm epoch when global surface temperatures varied between ∼5 and 8°C warmer than today, and CO₂concentration was ∼400–800 ppm. The narrowing/closing of the tropical ocean gateways and widening of high‐latitude gateways throughout the Miocene is likely responsible for the evolution of the ocean's overturning circulation to its modern structure, though the mechanisms remain unclear. Here, we investigate early and middle Miocene ocean circulation in an opportunistic climate model intercomparison (MioMIP1), using 14 simulations with different paleogeography, CO₂, and vegetation. The strength of the Southern Ocean‐driven Meridional Overturning Circulation (SOMOC) bottom cell is similar in the Miocene and Pre‐Industrial (PI) but dominates the Miocene global MOC due to weaker Northern Hemisphere overturning. The Miocene Atlantic MOC (AMOC) is weaker than PI in all the simulations (by 2–21 Sv), possibly due to its connection with an Arctic that is considerably fresher than today. Deep overturning in the North Pacific (PMOC) is present in three simulations (∼5–10 Sv), of which two have a weaker AMOC, and one has a stronger AMOC (compared to its PMOC). Surface freshwater fluxes control northern overturning such that the basin with the least freshwater gain has stronger overturning. While the orography, which impacts runoff direction (Pacific vs. Atlantic), has an inconsistent impact on northern overturning across simulations, overall, features associated with the early Miocene—such as a lower Tibetan Plateau, the Rocky Mountains, and a deeper Panama Seaway—seem to favor PMOC over AMOC. 

Abstract End of 21st‐century hydroclimate projections suggest an expansion of subtropical dry zones, with Mediterranean and Sahel regions becoming much drier. However, paleobotanical assemblage evidence from the middle Miocene (17‐12 Ma), suggests both regions were instead humid environments. Here we show that by modifying regional sea surface temperatures (SST) in an Earth System Model (CESM1.2) simulation of the middle Miocene, the increased ocean evaporation and integrated water vapor flux overrides any drying effects associated with warming‐induced land‐surface evaporation driven by atmospheric CO₂concentrations. These modifications markedly reduce the bias in the model‐data comparison for this period. A vegetation model (BIOME4) forced with simulated climatologies predicts both regions were dominated by mixed forest, which is largely consistent with the paleobotanical record. This study unveils the potential for wetter subtropical Mediterranean climates associated with warming, presenting an alternative scenario from future drying projections with localized SST warming governing regional climate change. 

Abstract As the last time period when concentrations were near 400 ppm, the Pliocene Epoch (5.33–2.58 Ma) is a useful paleoclimate target for understanding future climate change. Existing estimates of global warming and climate sensitivity during the Pliocene rely mainly on model simulations. To reconstruct Pliocene climate and incorporate paleoclimate observations, we use data assimilation to blend sea‐surface temperature (SST) proxies with model simulations from the Pliocene Modeling Intercomparison Project 2 and the Community Earth System Models. The resulting reconstruction, “plioDA,” suggests that the mid‐Pliocene (3.25 Ma) was warmer than previously thought (on average 4.1°C warmer than preindustrial, 95% CI = 3.0°C–5.3°C), leading to a higher estimate of climate sensitivity (4.8°C per doubling of , 90% CI = 2.6°C–9.9°C). In agreement with previous work, the tropical Pacific zonal SST gradient during the mid‐Pliocene was moderately reduced (°C, 95% CI = –0.4°C). However, this gradient was more reduced during the early Pliocene (4.75 Ma, -2.3°C, 95% CI = -3.9t to -1.1°C), a time period that is also warmer than the mid‐Pliocene (4.8°C above preindustrial, 95% CI = 3.6°C–6.2°C). PlioDA reconstructs a fresh North Pacific and salty North Atlantic, supporting Arctic gateway closure and contradicting the presence of Pacific Deep Water formation. Overall, plioDA updates our view of global and spatial climate change during the Pliocene, as well as raising questions about the state of ocean circulation and the drivers of differences between the early and mid‐Pliocene. 

Abstract Climate models require boundary condition information, such as vegetation and soil distributions because they influence the mean state climate, and feedbacks can significantly influence regional climate and climate sensitivity to CO₂forcing. Information about past distributions comes primarily from the paleobotanical record, which is often supplemented by a vegetation model to fill data gaps. For recent past periods such as the Pliocene, a quantitative suitability assessment of these vegetation model simulations is sufficient. However, the Miocene Climate Optimum spanning 16.9–14.7 Ma was the warmest period on Earth over the last ∼25 million years and models struggle to reproduce those conditions for the range of paleogeographies and CO₂concentrations tested, particularly at high latitudes. Here we bring together the Miocene modeling and data communities to update previous vegetation reconstructions used for climate modeling with a new regional approach that relaxes the requirement for a single model simulation to be used, blending instead simulations forced by different paleogeographies and CO₂concentrations. This ensures the simulated vegetation is first, and foremost, consistent with the paleorecord and provides a baseline for future comparisons. The reconstruction shows global increases in forest cover at all latitudes as compared to today and extensive C₃grasslands across the high northern latitudes. Data gaps at high latitudes are filled with vegetation models forced by higher CO₂concentrations than were required at lower latitudes consistent with the inability of current models to simulate Miocene high latitude warmth. 

Abstract The Miocene (23.03–5.33 Ma) is recognized as a period with close to modern‐day paleogeography, yet a much warmer climate. With large uncertainties in future hydroclimate projections, Miocene conditions illustrate a potential future analog for the Earth system. A recent opportunistic Miocene Model Intercomparison Project 1 (MioMIP1) focused on synthesizing published Miocene climate simulations and comparing them with available temperature reconstructions. Here, we build on this effort by analyzing the hydrological cycle response to Miocene forcings across early‐to‐middle (E2MMIO; 20.03–11.6 Ma) and middle‐to‐late Miocene (M2LMIO; 11.5–5.33 Ma) simulations with CO₂concentrations ranging from 200 to 850 ppm and providing a model‐data comparison against available precipitation reconstructions. We find global precipitation increases by ∼2.1 and 2.3% per degree of warming for E2MMIO and M2LMIO simulations, respectively. Models generally agree on a wetter than modern‐day tropics; mid and high‐latitude, however, do not agree on the sign of subtropical precipitation changes with warming. Global monsoon analysis suggests most monsoon regions, except the North American Monsoon, experience higher precipitation rates under warmer conditions. Model‐data comparison shows that mean annual precipitation is underestimated by the models regardless of CO₂concentration, particularly in the mid‐ to high‐latitudes. This suggests that the models may not be (a) resolving key processes driving the hydrological cycle response to Miocene boundary conditions and/or (b) other boundary conditions or processes not considered here are critical to reproducing Miocene hydroclimate. This study highlights the challenges in modeling and reconstructing the Miocene hydrological cycle and serves as a baseline for future coordinated MioMIP efforts. 

Characterized by similar-to-today CO2 (∼400 ppm) and surface temperatures approximately 3°–4°C warmer than the preindustrial, the mid-Pliocene warm period (mPWP) has often been used as an analog for modern CO2-driven climate change and as a constraint on the equilibrium climate sensitivity (ECS). However, model intercomparison studies suggest that non-CO2boundary conditions—such as changes in ice sheets—contribute substantially to the higher global mean temperatures and strongly shape the pattern of sea surface warming during the mPWP. Here, we employ a set of CESM2 simulations to quantify mPWP effective radiative forcings, study the role of ocean circulation changes in shaping the patterns of sea surface temperatures, and calculate radiative feedbacks during the mPWP. We find that the non-CO2boundary conditions of the mPWP, enhanced by changes in ocean circulation, contributed to larger high-latitude warming and less-stabilizing feedbacks relative to those induced by CO2alone. Accounting for differences in feedbacks between the mPWP and the modern (greenhouse gas–driven) climate provides stronger constraints on the high-end of modern-day ECS. However, a quantification of the forcing of non-CO₂boundary condition changes combined with the distinct radiative feedbacks that they induce suggests that Earth system sensitivity may be higher than previously estimated. 

Abstract Oxygen minimum zones (OMZs) play a critical role in global biogeochemical cycling and act as barriers to dispersal for marine organisms. OMZs are currently expanding and intensifying with climate change, however past distributions of OMZs are relatively unknown. Here we present evidence for widespread pelagic OMZs during the Pliocene (5.3-2.6 Ma), the most recent epoch with atmospheric CO 2 analogous to modern (~400-450 ppm). The global distribution of OMZ-affiliated planktic foraminifer, Globorotaloides hexagonus , and Earth System and Species Distribution Models show that the Indian Ocean, Eastern Equatorial Pacific, eastern South Pacific, and eastern North Atlantic all supported OMZs in the Pliocene, as today. By contrast, low-oxygen waters were reduced in the North Pacific and expanded in the North Atlantic in the Pliocene. This spatially explicit perspective reveals that a warmer world can support both regionally expanded and contracted OMZs, with intermediate water circulation as a key driver.